Kickxia |
Kickxia elatine |
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cancerwort, fluellin |
sharp leaf fluellin, sharp point fluellin, sharp-leaf cancerwort, sharp-leaf fluellen, sharppoint fluvellin |
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Habit | Herbs, annual [perennial]. | Plants 4–120 cm. |
Stems | decumbent or prostrate to erect [climbing], villous to glandular-hairy [glabrous]. |
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Leaves | cauline, alternate, sometimes opposite proximally; petiole present; blade oblong-ovate to orbiculate or cordate, not fleshy, not leathery, margins entire or dentate [lobed]. |
petiole 2–6 mm; blade: proximal 15–35 × 10–30 mm, base truncate or rounded to cuneate, distal 4–25 × 2–14 mm, base hastate to sagittate, villous with scattered glandular hairs. |
Inflorescences | axillary, flowers solitary; bracts absent. |
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Pedicels | present, spreading; bracteoles absent. |
10–22 mm. |
Flowers | bisexual; sepals 5, basally connate, equal, calyx radially symmetric, campanulate, lobes lanceolate [linear], villous; corolla yellow [white], often tinged blue to violet [red], bilaterally symmetric, bilabiate and personate, tubular, tube base not gibbous, spurred abaxially, lobes 5, abaxial 3, spreading, adaxial 2, erect, subequal, apex round; stamens 4, basally adnate to corolla, didynamous, included, filaments incurved, sparsely hairy or glabrous, anthers coherent, ciliate, pollen sacs oblong; staminode 0; ovary 2-locular, villous, locules subequal, placentation axile; stigma capitate. |
sepal lobes lanceolate, 3–6 × 1–2 mm, not accrescent, apex acuminate, villous; corolla tube 1–2 mm, palate yellow, inflated, hairy, spurs 5–6 mm, abaxial lobes yellow to violet, adaxial blue to violet, 2–3 mm; abaxial filaments 1.5–2 mm, adaxial 1–1.5 mm; style included, 1.3–1.5 mm; stigma straight. |
Fruits | capsules, dehiscence poricidal [indehiscent]. |
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Capsules | ovoid-globular, 3–5 mm. |
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Seeds | [6–]20–40[–60], dark brown to black, oblong-ovoid [reniform], wings absent. |
1–1.5 mm, cristate-tuberculate. |
x | = 9. |
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2n | = 36. |
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Kickxia |
Kickxia elatine |
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Phenology | Flowering May–Oct(–Dec). | |
Habitat | Gravelly or sandy disturbed sites, roadsides, stream banks, gravel bars, glades. | |
Elevation | 0–900 m. (0–3000 ft.) | |
Distribution |
Eurasia; n Africa; Atlantic Islands [Introduced in North America; introduced also in Mexico, West Indies, Central America, South America, Pacific Islands, Australia] |
AL; AR; CA; CT; DE; GA; IL; IN; KS; KY; LA; MA; MD; MI; MO; NC; NJ; NY; OH; OK; OR; PA; RI; SC; TN; TX; VA; WA; WI; WV; BC; ON; Eurasia; n Africa [Introduced in North America; introduced also in Mexico, Central America, South America, Atlantic Islands, Australia]
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Discussion | Species 9 or 46 (2 in the flora). Kickxia is characterized by personate, long-spurred corollas, coherent anthers, and poricidal capsule dehiscence. Early treatments proposed that Kickxia was congeneric with, or closely related to, Linaria (G. Bentham 1876; P. A. Munz 1926); later taxonomic treatments and phylogenetic analyses have supported generic rank for Kickxia (W. Rothmaler 1943; D. A. Sutton 1988; M. Ghebrehiwet 2001). Morphology- and molecular-based phylogenies support Anarrhinum Desfontaines (x = 10) as the most likely sister group to Kickxia (x = 9) and their basal position in Antirrhineae as the Anarrhinum clade (Ghebrehiwet et al. 2000; P. Vargas et al. 2004). Although usually considered in a broad sense (D. A. Sutton 1988), N. Yousefi et al. (2016) proposed, on the basis of ITS and trnl sequence data, that Kickxia sect. Valvatae (Wettstein) Janchen be recognized at the genus level. Kickxia then becomes a genus of nine species, with 37 being transferred to Nanorrhinum Betsche. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Kickxia elatine differs from K. spuria in leaf and calyx characters. Two subspecies of K. elatine have been described [subsp. elatine and subsp. crinita (Mabille) W. Greuter]; they are poorly defined (D. A. Sutton 1988; M. Ghebrehiwet 2001) and differentiated on the basis of indument density. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Key | ||
Source | FNA vol. 17, p. 25. | FNA vol. 17, p. 25. |
Parent taxa | ||
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Antirrhinum elatine | |
Name authority | Dumortier: Fl. Belg., 35. (1827) | (Linnaeus) Dumortier: Fl. Belg., 35. (1827) |
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