Kallstroemia maxima |
Zygophyllaceae |
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big caltrop |
caltrop family, creosote bush family |
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Habit | Herbs, annual. | Herbs, subshrubs, shrubs, or trees, annual or perennial, branching usually divaricate, growth sympodial, nodes angled or swollen, evergreen [deciduous], synoecious [dioecious]. | ||||||||||||||||||||
Stems | prostrate to decumbent, to 1(–1.5) m, sericeous and sparingly hirsute with white or yellow antrorse hairs, becoming glabrate. |
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Leaves | obovate in outline, 1–6 × 1.5–5 cm; stipules 3–5 × 1 mm; leaflets 6–8(–12), broadly oblong to elliptic, 5–29 × 3–14 mm, terminal pair usually largest, surfaces appressed-hirsute to glabrate, veins and margins sericeous. |
opposite or fascicled [alternate or on short lateral branches], palmately or even- [odd-]pinnately compound [simple]; stipules present; petiole present [absent]; blade often fleshy or coriaceous, margins entire; venation pinnate. |
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Inflorescences | pseudoaxillary [terminal], flowers solitary or in 2-flowered clusters [cymes]. |
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Pedicels | 10–50 mm in flower and fruit, at first shorter than subtending leaves, equaling them or longer in fruit, little thickened distally, straight or curved. |
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Flowers | 7–25 mm diam.; sepals persistent, ovate, 3–8 × 2–3 mm, as long as or little shorter than petals, in flower as long as style, in fruit clasping but not entirely covering mature fruit body and shorter than beak, only scarious margins becoming involute, hirsute; petals marcescent, usually ± 2-colored, basally usually white to yellow-green or green, rarely red (often brighter than distal portion), fading white to bright orange, distally white to yellow or pale orange, obovate, 5–12 × 4–10 mm; stamens as long as style; anthers yellow or red-orange, usually ovoid, rarely linear, 1 mm; ovary ovoid, 1 mm, usually glabrous, sometimes strigose at base, rarely to base of style; style cylindric but slightly conic basally, 2–3 mm, 2–3 times as long as ovary, glabrous; stigma terminal. |
bisexual [unisexual], usually regular, sometimes slightly irregular; perianth and androecium hypogynous; hypanthium absent; sepals 4–5, usually distinct, rarely connate basally; petals 4–5, distinct [rarely connate basally], often clawed, sometimes twisted; nectary usually present, extrastaminal and/or intrastaminal, rarely absent; stamens [5–](8–)10 in 2 whorls, outer usually opposite petals, often alternately unequal in length or sterile, distinct, free or adnate to petal bases, inserted on or proximal to nectary, frequently glandular or appendaged at base; anthers dehiscing by longitudinal slits; pistil 1, (2–)5-carpellate, ovary superior, (2–)5–10-locular; placentation axile [basal]; ovules (1–)2–10 per locule, anatropous; style 1; stigma 1. |
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Fruits | capsules, dehiscence septicidal or loculicidal, or schizocarps splitting into 5 or 10 mericarps. |
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Seeds | 1–5(–10) per locule. |
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Schizocarps | ovoid, 5–6 mm diam., usually glabrous, sometimes strigose at base or rarely to base of beak; beak cylindric, 3–7 mm, usually as long as fruit body, base widely conic, glabrous; mericarps 3–4 × 1 mm, abaxially tuberculate, cross-ridged, or slightly keeled, tubercles if present all rounded, less than 1 mm, sides pitted, adaxial edge angled. |
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Kallstroemia maxima |
Zygophyllaceae |
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Phenology | Flowering year-round. | |||||||||||||||||||||
Habitat | Weedy habitats, disturbed areas. | |||||||||||||||||||||
Elevation | 0–1400 m. (0–4600 ft.) | |||||||||||||||||||||
Distribution |
AL; FL; GA; SC; TX; Mexico; Central America; West Indies; n South America
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North America; Mexico; Central America; South America; West Indies; Eurasia; Africa; Atlantic Islands; Indian Ocean Islands; Pacific Islands; Australia; mostly tropical or subtropical regions; mainly in arid and semiarid areas |
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Discussion | The most widespread species of the genus, Kallstroemia maxima is usually found at lower elevations. The species probably has been taken to many places inadvertently by humans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 27, species ca. 240 (6 genera, 15 species in the flora). Zygophyllaceae are most closely related to Krameriaceae and the two families make up the isolated order Zygophyllales (Angiosperm Phylogeny Group 2003). A number grow in saline soils. Three species are cultivated in milder winter areas of the southeastern United States for their colorful flowers: the South American Bulnesia arborea (Jacquin) Engler and B. sarmientoi Lorentz ex Grisebach, both verawood, and the Caribbean Guaiacum officinale Linnaeus, lignum vitae. Guaiacum coulteri A. Gray, guayacán, from western Mexico and Guatemala, is grown in southern Arizona. Peganum, often placed in the Zygophyllaceae, is now recognized to be a member of the unrelated Nitrariaceae (Angiosperm Phylogeny Group). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 41. | FNA vol. 12, p. 28. | ||||||||||||||||||||
Parent taxa | Zygophyllaceae > Kallstroemia | |||||||||||||||||||||
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Subordinate taxa | ||||||||||||||||||||||
Synonyms | Tribulus maximus | |||||||||||||||||||||
Name authority | (Linnaeus) Hooker & Arnott: Bot. Beechey Voy., 282. (1838) | R. Brown | ||||||||||||||||||||
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