dubius rush, mariposa rush, questionable rush
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Herbs, perennial, rhizomatous, 3–7.5 dm. |
Herbs, perennial (rarely annual), rhizomatous. |
2–3 mm diam., not swollen. |
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erect, terete, 2–4 mm diam., smooth or rugulose. |
terete. |
1–2, pink to straw-colored, apex acute. |
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basal 1–2, cauline 1–2; auricles 1–4.9 mm, , apex rounded, membranaceous; blade green to straw-colored, terete, 20–30 cm × 1.5–4 mm, rugulose or smooth. |
basal and cauline: blade terete, nearly terete, or flattened, perfectly septate. |
terminal panicles of 25–66 heads, 7–13 cm, branches spreading; primary bract erect; heads 6–10-flowered, hemispheric to obovoid, 5–10 mm diam. |
terminal panicles or racemes of 2 to –many heads, a single terminal head, or rarely a cyme of heads, sympodial; bracts erect or ascending, flat, involute or terete; bracteoles absent. |
tepals straw-colored to brown, lanceolate, apex acuminate; outer tepals (2–)2.5–3.4 mm; inner tepals (2–)2.6–3.6 mm; stamens 6, anthers 1.5–2 times filament length. |
in multiflowered heads (rarely only 2– or 3 flowers per head). |
exserted, chestnut brown, 1-locular, subuloidsubulate, (2.4–)3–3.9 mm, apex tapering to subulate tip, valves separating at dehiscence. |
1- or 3-locular, beaked. |
obovoid, 0.3–0.4 mm, not tailed. |
tailed or not tailed. |
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= 20. |
= 40. |
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Fruiting early summer–late fall. |
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Montane meadows, riverbeds, canyons, aroyos |
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100–1600 m (300–5200 ft) |
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CA
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Worldwide |
Juncus dubius has rugulose stems and leaves throughout most of its range, but on the periphery (in Mariposa, San Diego, and Sonoma counties, California) the plants are smooth. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 80 (32 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Seeds tailed, 0.7–2.6 mm, including tails; seed body covered with whitish translucent veil. | → 2 |
1. Seeds not tailed, 0.3–0.7 mm; seed body clear yellow-brown. | → 7 |
2. Seeds 1.8–2.6 mm, including tails; stamens usually 6. | → 3 |
2. Seeds 0.7–1.9 mm, including tails; stamens usually 3. | → 4 |
3. Stems and leaves scabrous; auricles 1–7.5 mm | J. caesariensis |
3. Stems and leaves smooth; auricles 0.2–0.4 mm | J. trigonocarpus |
4. Seeds 1.1–1.9 mm; heads 5–50-flowered; inflorescence branches erect to ascending | J. canadensis |
4. Seeds 0.7–1.2 mm; heads 2–8-flowered or if to 20-flowered then proximal inflorescence branches widely spreading (J. subcaudatus). | → 5 |
5. Outer tepals obtuse to nearly acute | J. brachycephalus |
5. Outer tepals acuminate to acute. | → 6 |
6. Proximal inflorescence branches widely spreading | J. subcaudatus |
6. Proximal inflorescence branches erect | J. brevicaudatus |
7. Flowers 1–2(–4) at each node, not in heads; capsules fertile only below middle. | → 8 |
7. Flowers in heads of 3–60; capsules fertile throughout or only below middle. | → 9 |
8. Culms erect, 0.3–7 dm, not forming mats | J. pelocarpus |
8. Culms decumbent, 0.1–1 dm, forming mats | J. subtilis |
9. Heads spheric or nearly spheric, 15–60-flowered | → 10 |
9. Heads obconic to hemispheric, 3–15-flowered. | → 23 |
| → 18 |
| → 11 |
11. Plants cespitose; tepals lanceolate | J. acuminatus |
11. Plants rhizomatous; tepals lance-subulate. | → 12 |
| J. brachycarpus |
12. Capsule exserted to slightly included. | → 13 |
13. Capsules remaining united at apex after dehiscence. | → 14 |
13. Capsule valves separating at apex during dehiscence. | → 16 |
14. Leaves laterally compressed, 3–5 mm diam. | J. validus var. fascinatus |
14. Leaves terete, 1–4 mm diam. | → 15 |
15. Heads sometimes spheric, usually lobed; tepals green to stramineous, nearly equal; uppermost cauline leaf blade (below inflorescence bract) equaling or longer than its sheath | J. scirpoides |
15. Heads spheric; stramineous to reddish brown, inner tepals shorter than outer; uppermost cauline leaf blade much shorter than its sheath | J. megacephalus |
16. Tepals 4–5 mm; leaves laterally compressed | J. validus var. validus |
16. Tepals 2.9–4 mm; leaves terete. | → 17 |
17. Capsules 3.5–5 mm, exserted; culms 0.4–3 dm | J. nodosus |
17. Capsules 3–3.5 mm; equaling perianth or slightly shorter; culms 2.5–8.5 dm | J. bolanderi |
18. Tepals lanceolate to lance-ovate, light green to light pink or white or dark purplish brown; capsule ovoid to obovoid, 1.9–3.5 mm; heads 1–2 per inflorescence (15–50 for J. pervetus). | → 19 |
18. Tepals lance-subulate, green to stramineous; capsule lance-subulate, 3.2–6.5 mm; heads 1–23 per inflorescence. | → 21 |
| J. pervetus |
| → 20 |
20. Tepals dark purplish brown to black; anthers 1/4 to equal filament length | J. mertensianus |
20. Tepals light green, or light pink to white; anthers 2–3 times length of filaments | J. chlorocephalus |
21. Outer tepals 2.4–4.1 mm, equaling inner tepals; auricles 0.5–1.7 mm | J. nodosus |
21. Outer tepals 4–6 mm, outer and inner tepals unequal in length; auricles 1–4 mm. | → 22 |
22. Outer tepals longer than inner; capsule 4.3–5.7 mm | J. torreyi |
22. Outer tepals shorter than inner; capsules 5.2–6.5 mm | J. texanus |
| → 24 |
| → 31 |
24. Stems creeping or floating, capillary; filiform, submersed leaves often formed before flowering; inflorescences with 1–9 heads. | → 25 |
24. Stems erect, not floating; filiform leaves absent; inflorescences with 3–200 heads. | → 26 |
25. Capsule rounded at apex, 2.5–4 mm; tepals 2–3.6 mm, obtuse to acute | J. bulbosus |
25. Capsule acute to acuminate at apex, 3.5–6.1 mm; tepals 2.8–5.5 mm, acute to acuminate-subulate | J. supiniformis |
26. Capsule much longer than perianth, 1.3–2 times perianth length. | → 27 |
26. Capsule equaling or just exceeding perianth. | → 29 |
27. Capsules beaked; plants often decumbent or matted | J. supiniformis |
27. Capsules not beaked; plants erect. | → 28 |
28. Capsule 1/3 longer than perianth, 2.8–4.2 mm | J. debilis |
28. Capsule twice as long as perianth, 4–5.2 mm | J. diffusissimus |
29. Heads 5–50; tepals 2.6–3.5 mm, nearly equal; capsules 2.8–3.5 mm | J. acuminatus |
29. Heads 30–250; tepals 1.7–2.9 mm, inner tepals shorter than outer tepals; capsules 1.9–2.9 mm. | → 30 |
30. Leaf blade with prominent and conspicuous ringlike bands at position of cross partitions; roots without terminal tubers; inner tepals 1.7–2.1 mm | J. nodatus |
30. Leaf blade with faint ringlike bands at position of septa; roots often with terminal tubers; inner tepals 2.4–2.8 mm | J. elliottii |
31. Stems creeping or floating, capillary; filiform, submersed leaves often formed before flowering; inflorescences with 1–9 heads. | → 32 |
31. Stems erect; filiform submersed leaves not formed except in J. militaris; heads 1–66. | → 34 |
32. Auricles 0.8–2.1 mm; capsules 3.5–6.1 mm | J. supiniformis |
32. Auricles 0.4–1 mm; capsules 2.5–4 mm. | → 33 |
33. Capsules exserted about 1 mm beyond perianth; apex acute below beak; outer tepals acute to acuminate | J. articulatus |
33. Capsules as long as tepals or exserted to 0.5 mm beyond tepals, apex obtuse below apex; outer tepals acute to obtuse | J. bulbosus |
34. Proximal culm leaf overtopping inflorescences, distal culm leaf usually inflated, bladeless sheath; often developing filiform (hairlike) leaves off rhizome in running water | J. militaris |
34. Proximal culm leaf shorter than inflorescence, distal culm leaf usually with blade, sheath not inflated; not developing filiform leaves off rhizome. | → 35 |
35. Tepals 1.6–3 mm; apex obtuse to acuminate, usually not subulate mucronate; anthers shorter than to equaling filaments. | → 36 |
35. Tepals 2.3–6.2 mm; apex usually acuminate with subulate mucro (acute in J. acutiflorus); anthers equaling or longer than filaments. | → 37 |
36. Inner tepals obtuse; inflorescence stiffly erect | J. alpinoarticulatus |
36. Inner tepals acuminate; inflorescence spreading | J. articulatus |
37. Capsule abruptly contracted at apex, usually not exserted | J. nevadensis |
37. Capsule narrowed to beak, exserted at least slightly. | → 38 |
38. Heads 2–9; tepals 2.8–5.5 mm | J. supiniformis |
38. Heads 25–66; tepals 2.5–3.5 mm. | → 39 |
39. Tepals nearly equal; heads with 6–10 flowers | J. dubius |
39. Tepals unequal, outer shorter than inner; heads with 5–8 flowers | J. acutiflorus |
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FNA vol. 22. |
FNA vol. 22. |
Juncaceae > Juncus > subg. Septati |
Juncaceae > Juncus |
J. acuminatus, J. acutiflorus, J. acutus, J. alpinoarticulatus, J. anthelatus, J. arcticus, J. articulatus, J. biglumis, J. bolanderi, J. brachycarpus, J. brachycephalus, J. brachyphyllus, J. brevicaudatus, J. bryoides, J. bufonius, J. bulbosus, J. caesariensis, J. canadensis, J. capillaris, J. capitatus, J. castaneus, J. chlorocephalus, J. compressus, J. confusus, J. cooperi, J. coriaceus, J. covillei, J. debilis, J. dichotomus, J. diffusissimus, J. drummondii, J. dudleyi, J. effusus, J. elliottii, J. ensifolius, J. falcatus, J. filiformis, J. filipendulus, J. georgianus, J. gerardii, J. greenei, J. gymnocarpus, J. hallii, J. hemiendytus, J. howellii, J. inflexus, J. interior, J. kelloggii, J. leiospermus, J. lesueurii, J. longistylis, J. luciensis, J. macrandrus, J. macrophyllus, J. marginatus, J. maritimus, J. megacephalus, J. mertensianus, J. militaris, J. nevadensis, J. nodatus, J. nodosus, J. occidentalis, J. orthophyllus, J. oxymeris, J. parryi, J. patens, J. pelocarpus, J. pervetus, J. phaeocephalus, J. polycephalus, J. regelii, J. repens, J. roemerianus, J. scirpoides, J. secundus, J. squarrosus, J. stygius, J. subcaudatus, J. subtilis, J. supiniformis, J. tenuis, J. texanus, J. textilis, J. tiehmii, J. torreyi, J. trifidus, J. triformis, J. triglumis, J. trigonocarpus, J. uncialis, J. validus, J. vaseyi, J. xiphioides |
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J. acuminatus, J. acutiflorus, J. alpinoarticulatus, J. articulatus, J. bolanderi, J. brachycarpus, J. brachycephalus, J. brevicaudatus, J. bulbosus, J. caesariensis, J. canadensis, J. chlorocephalus, J. debilis, J. diffusissimus, J. dubius, J. elliottii, J. megacephalus, J. mertensianus, J. militaris, J. nevadensis, J. nodatus, J. nodosus, J. pelocarpus, J. pervetus, J. scirpoides, J. subcaudatus, J. subtilis, J. supiniformis, J. texanus, J. torreyi, J. trigonocarpus, J. validus var. fascinatus, J. validus var. validus |
J. rugulosus |
J. section Septati |
Engelmann |
Buchenau: Abhandlungen herausgegeben vom naturwissenschaftlichen Vereine zu Bremen 4: 406. (1875) |
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