FNA: Juncus bufonius vs. Juncus nevadensis

	Juncus bufonius	Juncus nevadensis
	common toad rush, toad rush	Nevada rush, Sierra rush
Habit	Herbs, annual, cespitose, 0.5–4 dm.	Herbs, perennial, rhizomatous, 0.5–7 dm.
Rhizomes		1 mm diam., not swollen.
Culms	1–many, occasionally becoming decumbent.	erect, terete, 1.5–3 mm diam., smooth.
Cataphylls	0–2.	0–1, maroon or straw-colored, apex acute.
Leaves	basal and cauline; auricles rudimentary or absent; blade flat, 3–13 cm × 0.3–1.1 mm.	basal 1–3, cauline 1–2; auricles 1–3.2 mm, apex rounded to acute, membranaceous; blade green, laterally flattened, 1.5–31 cm × 0.5–2.2 mm.
Inflorescences	loose and diffuse or less often compact, usually at least ½1/2 total height of plant; primary bract shorter than inflorescence.	terminal panicles of 2–11 heads, 2–18 cm, branches erect to spreading; primary bract erect; heads 3–11-flowered, hemispheric to obpyramidal (rarely nearly spheric), 5–14 mm diam.
Flowers	bracteoles 2; tepals greenish, lanceolate, 3.8–7(–8.5) mm; inner series slightly shorter, apex sometimes obtuse; stamens 3–6, filaments (0.7–)1–1.8 mm, anthers 0.3–0.8 mm; style 0.1–0.2 mm.	tepals dark brown to white, lanceolate, apex acute to acuminate, mucro subulate; outer tepals 2.8–6.2 mm; inner tepals 2.4–6 mm; stamens 6, anthers 1–2 times filament length.
Capsules	tan to reddish brown, 3-locular, ellipsoid to narrowly so, slightly truncate, 2.7–4 × 1–1.5 mm, sometimes exceeding inner tepals but usually not outer series.	included to slightly exserted, chestnut brown, ellipsoid, 2.3–3.7 mm, abruptly contracted to apex, apex acute proximal to beak, valves separating at dehiscence.
Seeds	yellowish, widely ellipsoid to ovoid, 0.26–0.49, not tailed.	ellipsoid, 0.4–0.5 mm, not tailed.
2n	= 27–37, 58–81, 108–115.

	Juncus bufonius	Juncus nevadensis
Phenology	Flowering and fruiting spring–early fall.	Fruiting early summer–fall.
Habitat	Moist soils in meadows, along lakeshores or stream banks, ditches, or roadsides, especially frequent in drawdown areas, usually in open sites and often becoming weedy	Wet banks along margins of streams and lakes, marshes, bogs, sloughs, and wet or boggy prairies
Elevation		0–2600 m (0–8500 ft)
Distribution	AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; Greenland; nearly worldwide [WildflowerSearch map] [BONAP county map]	AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; SK [WildflowerSearch map] [BONAP county map]
Discussion	Nearly worldwide, Juncus bufonius is found essentially throughout North America except north of the Alaskan and Canadian tTaiga. Juncus bufonius is a highly polymorphic complex that is poorly understood systematically. Insufficient evidence exists upon which to base the segregation of the plethora of taxa that have been recognized out of this group in the past. (Discussion copyrighted by Flora of North America; reprinted with permission.)	This variable species has been divided into five separate taxa in the past (F. J. Hermann 1964; C. L. Hitchcock et al. 1955–1969, vol. 1), but more recently, most of the variants have not been separated (A. Cronquist et al. 1972+, vol. 6). The Oregon coastal endemic, Juncus nevadensis var. inventus L. F. Henderson) C. L. Hitchcock, is at one extreme of the variation, having only a single head, fairly dark brown tepals 3.5–4.5 mm, anthers only slightly longer than the filaments, and a distinct habitat. The J. mertensianus var. columbianus segregate, however, approaches var. inventus in number of heads, and the other characters are so highly variable in the complex, they cannot be used alone to separate this variety. Therefore, we are not recognizing any infraspecific taxa at this time. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 22.	FNA vol. 22.
Parent taxa	Juncaceae > Juncus > subg. Poiophylli	Juncaceae > Juncus > subg. Septati
Sibling taxa	J. acuminatus, J. acutiflorus, J. acutus, J. alpinoarticulatus, J. anthelatus, J. arcticus, J. articulatus, J. biglumis, J. bolanderi, J. brachycarpus, J. brachycephalus, J. brachyphyllus, J. brevicaudatus, J. bryoides, J. bulbosus, J. caesariensis, J. canadensis, J. capillaris, J. capitatus, J. castaneus, J. chlorocephalus, J. compressus, J. confusus, J. cooperi, J. coriaceus, J. covillei, J. debilis, J. dichotomus, J. diffusissimus, J. drummondii, J. dubius, J. dudleyi, J. effusus, J. elliottii, J. ensifolius, J. falcatus, J. filiformis, J. filipendulus, J. georgianus, J. gerardii, J. greenei, J. gymnocarpus, J. hallii, J. hemiendytus, J. howellii, J. inflexus, J. interior, J. kelloggii, J. leiospermus, J. lesueurii, J. longistylis, J. luciensis, J. macrandrus, J. macrophyllus, J. marginatus, J. maritimus, J. megacephalus, J. mertensianus, J. militaris, J. nevadensis, J. nodatus, J. nodosus, J. occidentalis, J. orthophyllus, J. oxymeris, J. parryi, J. patens, J. pelocarpus, J. pervetus, J. phaeocephalus, J. polycephalus, J. regelii, J. repens, J. roemerianus, J. scirpoides, J. secundus, J. squarrosus, J. stygius, J. subcaudatus, J. subtilis, J. supiniformis, J. tenuis, J. texanus, J. textilis, J. tiehmii, J. torreyi, J. trifidus, J. triformis, J. triglumis, J. trigonocarpus, J. uncialis, J. validus, J. vaseyi, J. xiphioides	J. acuminatus, J. acutiflorus, J. acutus, J. alpinoarticulatus, J. anthelatus, J. arcticus, J. articulatus, J. biglumis, J. bolanderi, J. brachycarpus, J. brachycephalus, J. brachyphyllus, J. brevicaudatus, J. bryoides, J. bufonius, J. bulbosus, J. caesariensis, J. canadensis, J. capillaris, J. capitatus, J. castaneus, J. chlorocephalus, J. compressus, J. confusus, J. cooperi, J. coriaceus, J. covillei, J. debilis, J. dichotomus, J. diffusissimus, J. drummondii, J. dubius, J. dudleyi, J. effusus, J. elliottii, J. ensifolius, J. falcatus, J. filiformis, J. filipendulus, J. georgianus, J. gerardii, J. greenei, J. gymnocarpus, J. hallii, J. hemiendytus, J. howellii, J. inflexus, J. interior, J. kelloggii, J. leiospermus, J. lesueurii, J. longistylis, J. luciensis, J. macrandrus, J. macrophyllus, J. marginatus, J. maritimus, J. megacephalus, J. mertensianus, J. militaris, J. nodatus, J. nodosus, J. occidentalis, J. orthophyllus, J. oxymeris, J. parryi, J. patens, J. pelocarpus, J. pervetus, J. phaeocephalus, J. polycephalus, J. regelii, J. repens, J. roemerianus, J. scirpoides, J. secundus, J. squarrosus, J. stygius, J. subcaudatus, J. subtilis, J. supiniformis, J. tenuis, J. texanus, J. textilis, J. tiehmii, J. torreyi, J. trifidus, J. triformis, J. triglumis, J. trigonocarpus, J. uncialis, J. validus, J. vaseyi, J. xiphioides
Synonyms	J. bufonius var. congestus, J. bufonius var. halophilus, J. bufonius var. hybridus, J. bufonius var. occidentalis, J. bufonius var. ranarius, J. congestus, J. ranarius	J. badius, J. columbianus, J. mertensianuss subsp. gracilis, J. mertensianus var. badius, J. mertensianus var. columbianus, J. mertensianus var. gracilis, J. mertensianus var. suksdorfii, J. nevadensis var. badius, J. nevadensis var. columbianus, J. nevadensis var. inventus, J. phaeocephalus var. gracilis, J. suksdorfii, J. truncatus
Name authority	Linnaeus: Sp. Pl. 1: 328. (1753)	S. Watson: Proceedings of the American Academy of Arts and Sciences 14: 303. (1879)
Web links	Local floras: BC, CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, Go Botany, IL Wildflowers, KS Wildflowers, LA Plants, MD Biodiversity, MI Flora, MN Wildflowers, MO Plants, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: BC, CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Juncus nevadensis

common toad rush, toad rush

Nevada rush, Sierra rush

Habit

Herbs, annual, cespitose, 0.5–4 dm.

Herbs, perennial, rhizomatous, 0.5–7 dm.

Rhizomes

1 mm diam., not swollen.

Culms

1–many, occasionally becoming decumbent.

erect, terete, 1.5–3 mm diam., smooth.

Cataphylls

0–2.

0–1, maroon or straw-colored, apex acute.

Leaves

basal and cauline;

auricles rudimentary or absent;

blade flat, 3–13 cm × 0.3–1.1 mm.

basal 1–3, cauline 1–2;

auricles 1–3.2 mm, apex rounded to acute, membranaceous;

blade green, laterally flattened, 1.5–31 cm × 0.5–2.2 mm.

Inflorescences

loose and diffuse or less often compact, usually at least ½1/2 total height of plant;

primary bract shorter than inflorescence.

terminal panicles of 2–11 heads, 2–18 cm, branches erect to spreading;

primary bract erect;

heads 3–11-flowered, hemispheric to obpyramidal (rarely nearly spheric), 5–14 mm diam.

Flowers

bracteoles 2;

tepals greenish, lanceolate, 3.8–7(–8.5) mm;

inner series slightly shorter, apex sometimes obtuse;

stamens 3–6, filaments (0.7–)1–1.8 mm, anthers 0.3–0.8 mm;

style 0.1–0.2 mm.

tepals dark brown to white, lanceolate, apex acute to acuminate, mucro subulate;

outer tepals 2.8–6.2 mm;

inner tepals 2.4–6 mm;

stamens 6, anthers 1–2 times filament length.

Capsules

tan to reddish brown, 3-locular, ellipsoid to narrowly so, slightly truncate, 2.7–4 × 1–1.5 mm, sometimes exceeding inner tepals but usually not outer series.

included to slightly exserted, chestnut brown, ellipsoid, 2.3–3.7 mm, abruptly contracted to apex, apex acute proximal to beak, valves separating at dehiscence.

Seeds

yellowish, widely ellipsoid to ovoid, 0.26–0.49, not tailed.

ellipsoid, 0.4–0.5 mm, not tailed.

= 27–37, 58–81, 108–115.

Juncus nevadensis

Phenology

Flowering and fruiting spring–early fall.

Fruiting early summer–fall.

Habitat

Moist soils in meadows, along lakeshores or stream banks, ditches, or roadsides, especially frequent in drawdown areas, usually in open sites and often becoming weedy

Wet banks along margins of streams and lakes, marshes, bogs, sloughs, and wet or boggy prairies

Elevation

0–2600 m (0–8500 ft)

Distribution

AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; Greenland; nearly worldwide

[WildflowerSearch map]

[BONAP county map]

AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; SK

[WildflowerSearch map]

[BONAP county map]

Discussion

Nearly worldwide, Juncus bufonius is found essentially throughout North America except north of the Alaskan and Canadian tTaiga. Juncus bufonius is a highly polymorphic complex that is poorly understood systematically. Insufficient evidence exists upon which to base the segregation of the plethora of taxa that have been recognized out of this group in the past.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

This variable species has been divided into five separate taxa in the past (F. J. Hermann 1964; C. L. Hitchcock et al. 1955–1969, vol. 1), but more recently, most of the variants have not been separated (A. Cronquist et al. 1972+, vol. 6). The Oregon coastal endemic, Juncus nevadensis var. inventus L. F. Henderson) C. L. Hitchcock, is at one extreme of the variation, having only a single head, fairly dark brown tepals 3.5–4.5 mm, anthers only slightly longer than the filaments, and a distinct habitat. The J. mertensianus var. columbianus segregate, however, approaches var. inventus in number of heads, and the other characters are so highly variable in the complex, they cannot be used alone to separate this variety. Therefore, we are not recognizing any infraspecific taxa at this time.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 22.

Parent taxa

Juncaceae > Juncus > subg. Poiophylli

Juncaceae > Juncus > subg. Septati

Sibling taxa

J. acuminatus, J. acutiflorus, J. acutus, J. alpinoarticulatus, J. anthelatus, J. arcticus, J. articulatus, J. biglumis, J. bolanderi, J. brachycarpus, J. brachycephalus, J. brachyphyllus, J. brevicaudatus, J. bryoides, J. bulbosus, J. caesariensis, J. canadensis, J. capillaris, J. capitatus, J. castaneus, J. chlorocephalus, J. compressus, J. confusus, J. cooperi, J. coriaceus, J. covillei, J. debilis, J. dichotomus, J. diffusissimus, J. drummondii, J. dubius, J. dudleyi, J. effusus, J. elliottii, J. ensifolius, J. falcatus, J. filiformis, J. filipendulus, J. georgianus, J. gerardii, J. greenei, J. gymnocarpus, J. hallii, J. hemiendytus, J. howellii, J. inflexus, J. interior, J. kelloggii, J. leiospermus, J. lesueurii, J. longistylis, J. luciensis, J. macrandrus, J. macrophyllus, J. marginatus, J. maritimus, J. megacephalus, J. mertensianus, J. militaris, J. nevadensis, J. nodatus, J. nodosus, J. occidentalis, J. orthophyllus, J. oxymeris, J. parryi, J. patens, J. pelocarpus, J. pervetus, J. phaeocephalus, J. polycephalus, J. regelii, J. repens, J. roemerianus, J. scirpoides, J. secundus, J. squarrosus, J. stygius, J. subcaudatus, J. subtilis, J. supiniformis, J. tenuis, J. texanus, J. textilis, J. tiehmii, J. torreyi, J. trifidus, J. triformis, J. triglumis, J. trigonocarpus, J. uncialis, J. validus, J. vaseyi, J. xiphioides

J. acuminatus, J. acutiflorus, J. acutus, J. alpinoarticulatus, J. anthelatus, J. arcticus, J. articulatus, J. biglumis, J. bolanderi, J. brachycarpus, J. brachycephalus, J. brachyphyllus, J. brevicaudatus, J. bryoides, J. bufonius, J. bulbosus, J. caesariensis, J. canadensis, J. capillaris, J. capitatus, J. castaneus, J. chlorocephalus, J. compressus, J. confusus, J. cooperi, J. coriaceus, J. covillei, J. debilis, J. dichotomus, J. diffusissimus, J. drummondii, J. dubius, J. dudleyi, J. effusus, J. elliottii, J. ensifolius, J. falcatus, J. filiformis, J. filipendulus, J. georgianus, J. gerardii, J. greenei, J. gymnocarpus, J. hallii, J. hemiendytus, J. howellii, J. inflexus, J. interior, J. kelloggii, J. leiospermus, J. lesueurii, J. longistylis, J. luciensis, J. macrandrus, J. macrophyllus, J. marginatus, J. maritimus, J. megacephalus, J. mertensianus, J. militaris, J. nodatus, J. nodosus, J. occidentalis, J. orthophyllus, J. oxymeris, J. parryi, J. patens, J. pelocarpus, J. pervetus, J. phaeocephalus, J. polycephalus, J. regelii, J. repens, J. roemerianus, J. scirpoides, J. secundus, J. squarrosus, J. stygius, J. subcaudatus, J. subtilis, J. supiniformis, J. tenuis, J. texanus, J. textilis, J. tiehmii, J. torreyi, J. trifidus, J. triformis, J. triglumis, J. trigonocarpus, J. uncialis, J. validus, J. vaseyi, J. xiphioides

Synonyms

J. bufonius var. congestus, J. bufonius var. halophilus, J. bufonius var. hybridus, J. bufonius var. occidentalis, J. bufonius var. ranarius, J. congestus, J. ranarius

J. badius, J. columbianus, J. mertensianuss subsp. gracilis, J. mertensianus var. badius, J. mertensianus var. columbianus, J. mertensianus var. gracilis, J. mertensianus var. suksdorfii, J. nevadensis var. badius, J. nevadensis var. columbianus, J. nevadensis var. inventus, J. phaeocephalus var. gracilis, J. suksdorfii, J. truncatus

Name authority

Linnaeus: Sp. Pl. 1: 328. (1753)

S. Watson: Proceedings of the American Academy of Arts and Sciences 14: 303. (1879)

Web links

Local floras: BC, CA, OR, WA
Local Web sites: CalFlora, CalPhotos, Flora NW, Go Botany, IL Wildflowers, KS Wildflowers, LA Plants, MD Biodiversity, MI Flora, MN Wildflowers, MO Plants, PNW Herbaria
WildflowerSearch
iNaturalist (observations)
USDA Plants Database
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Local floras: BC, CA, OR, WA
Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria
WildflowerSearch
iNaturalist (observations)
USDA Plants Database
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Flora of North America species comparison

Juncus bufonius

Juncus nevadensis

Juncus bufonius

Juncus nevadensis