FNA: Juncus bufonius vs. Juncus leiospermus

	Juncus bufonius	Juncus leiospermus
	common toad rush, toad rush	dwarf rush, red bluff dwarf rush, smooth-seed rush
Habit	Herbs, annual, cespitose, 0.5–4 dm.	Herbs, annual, cespitose, 0.2–1.2 dm.
Culms	1–many, occasionally becoming decumbent.	to 100, (0.2–)0.4–0.8 mm diam..
Cataphylls	0–2.
Leaves	basal and cauline; auricles rudimentary or absent; blade flat, 3–13 cm × 0.3–1.1 mm.	1/3–3/4 height of plant.
Inflorescences	loose and diffuse or less often compact, usually at least ½1/2 total height of plant; primary bract shorter than inflorescence.	terminal clusters, each with 1–7 flowers; bracts subtending inflorescence 2–8, round to acutely ovate, inconspicuous, 0.7–2.4 mm, membranous.
Flowers	bracteoles 2; tepals greenish, lanceolate, 3.8–7(–8.5) mm; inner series slightly shorter, apex sometimes obtuse; stamens 3–6, filaments (0.7–)1–1.8 mm, anthers 0.3–0.8 mm; style 0.1–0.2 mm.	tepals reddish to black, narrowly lanceolate to oblong, 2–4.6 × 0.5–1.4 mm; inner series usually slightly longer than outer, apex obtuse to acuminate; stamens 3, filaments 0.5–1.4 mm, anthers 1–3.2 mm; style persistent, 1.4–4 mm, stigma 1.3–4 mm.
Capsules	tan to reddish brown, 3-locular, ellipsoid to narrowly so, slightly truncate, 2.7–4 × 1–1.5 mm, sometimes exceeding inner tepals but usually not outer series.	reddish, 3-locular, globose to ellipsoid or oblong, 2–4.5 × 1–3 mm, nearly equal to sepals.
Seeds	yellowish, widely ellipsoid to ovoid, 0.26–0.49, not tailed.	ovoid, 0.35–0.45 mm.
n		= 16.
2n	= 27–37, 58–81, 108–115.

	Juncus bufonius	Juncus leiospermus
Phenology	Flowering and fruiting spring–early fall.	Flowering and fruiting in spring.
Habitat	Moist soils in meadows, along lakeshores or stream banks, ditches, or roadsides, especially frequent in drawdown areas, usually in open sites and often becoming weedy	Margins of vernal pools
Elevation		0–500 m (0–1600 ft)
Distribution	AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; Greenland; nearly worldwide [WildflowerSearch map] [BONAP county map]	CA [WildflowerSearch map] [BONAP county map]
Discussion	Nearly worldwide, Juncus bufonius is found essentially throughout North America except north of the Alaskan and Canadian tTaiga. Juncus bufonius is a highly polymorphic complex that is poorly understood systematically. Insufficient evidence exists upon which to base the segregation of the plethora of taxa that have been recognized out of this group in the past. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Plants from Butte and Calaveras Ccounties, California, tend to be smaller and consistently develop only one flower per culm; they may be recognized as Juncus leiospermus var. ahartii Ertter. Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 22.	FNA vol. 22.
Parent taxa	Juncaceae > Juncus > subg. Poiophylli	Juncaceae > Juncus > subg. Graminifolii
Sibling taxa	J. acuminatus, J. acutiflorus, J. acutus, J. alpinoarticulatus, J. anthelatus, J. arcticus, J. articulatus, J. biglumis, J. bolanderi, J. brachycarpus, J. brachycephalus, J. brachyphyllus, J. brevicaudatus, J. bryoides, J. bulbosus, J. caesariensis, J. canadensis, J. capillaris, J. capitatus, J. castaneus, J. chlorocephalus, J. compressus, J. confusus, J. cooperi, J. coriaceus, J. covillei, J. debilis, J. dichotomus, J. diffusissimus, J. drummondii, J. dubius, J. dudleyi, J. effusus, J. elliottii, J. ensifolius, J. falcatus, J. filiformis, J. filipendulus, J. georgianus, J. gerardii, J. greenei, J. gymnocarpus, J. hallii, J. hemiendytus, J. howellii, J. inflexus, J. interior, J. kelloggii, J. leiospermus, J. lesueurii, J. longistylis, J. luciensis, J. macrandrus, J. macrophyllus, J. marginatus, J. maritimus, J. megacephalus, J. mertensianus, J. militaris, J. nevadensis, J. nodatus, J. nodosus, J. occidentalis, J. orthophyllus, J. oxymeris, J. parryi, J. patens, J. pelocarpus, J. pervetus, J. phaeocephalus, J. polycephalus, J. regelii, J. repens, J. roemerianus, J. scirpoides, J. secundus, J. squarrosus, J. stygius, J. subcaudatus, J. subtilis, J. supiniformis, J. tenuis, J. texanus, J. textilis, J. tiehmii, J. torreyi, J. trifidus, J. triformis, J. triglumis, J. trigonocarpus, J. uncialis, J. validus, J. vaseyi, J. xiphioides	J. acuminatus, J. acutiflorus, J. acutus, J. alpinoarticulatus, J. anthelatus, J. arcticus, J. articulatus, J. biglumis, J. bolanderi, J. brachycarpus, J. brachycephalus, J. brachyphyllus, J. brevicaudatus, J. bryoides, J. bufonius, J. bulbosus, J. caesariensis, J. canadensis, J. capillaris, J. capitatus, J. castaneus, J. chlorocephalus, J. compressus, J. confusus, J. cooperi, J. coriaceus, J. covillei, J. debilis, J. dichotomus, J. diffusissimus, J. drummondii, J. dubius, J. dudleyi, J. effusus, J. elliottii, J. ensifolius, J. falcatus, J. filiformis, J. filipendulus, J. georgianus, J. gerardii, J. greenei, J. gymnocarpus, J. hallii, J. hemiendytus, J. howellii, J. inflexus, J. interior, J. kelloggii, J. lesueurii, J. longistylis, J. luciensis, J. macrandrus, J. macrophyllus, J. marginatus, J. maritimus, J. megacephalus, J. mertensianus, J. militaris, J. nevadensis, J. nodatus, J. nodosus, J. occidentalis, J. orthophyllus, J. oxymeris, J. parryi, J. patens, J. pelocarpus, J. pervetus, J. phaeocephalus, J. polycephalus, J. regelii, J. repens, J. roemerianus, J. scirpoides, J. secundus, J. squarrosus, J. stygius, J. subcaudatus, J. subtilis, J. supiniformis, J. tenuis, J. texanus, J. textilis, J. tiehmii, J. torreyi, J. trifidus, J. triformis, J. triglumis, J. trigonocarpus, J. uncialis, J. validus, J. vaseyi, J. xiphioides
Synonyms	J. bufonius var. congestus, J. bufonius var. halophilus, J. bufonius var. hybridus, J. bufonius var. occidentalis, J. bufonius var. ranarius, J. congestus, J. ranarius	J. leiospermus var. ahartii
Name authority	Linnaeus: Sp. Pl. 1: 328. (1753)	F. J. Hermann: Leaflets of Western Botany 5: 113. (1948)
Web links	Local floras: BC, CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, Go Botany, IL Wildflowers, KS Wildflowers, LA Plants, MD Biodiversity, MI Flora, MN Wildflowers, MO Plants, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Juncus leiospermus

common toad rush, toad rush

dwarf rush, red bluff dwarf rush, smooth-seed rush

Habit

Herbs, annual, cespitose, 0.5–4 dm.

Herbs, annual, cespitose, 0.2–1.2 dm.

Culms

1–many, occasionally becoming decumbent.

to 100, (0.2–)0.4–0.8 mm diam..

Cataphylls

0–2.

Leaves

basal and cauline;

auricles rudimentary or absent;

blade flat, 3–13 cm × 0.3–1.1 mm.

1/3–3/4 height of plant.

Inflorescences

loose and diffuse or less often compact, usually at least ½1/2 total height of plant;

primary bract shorter than inflorescence.

terminal clusters, each with 1–7 flowers;

bracts subtending inflorescence 2–8, round to acutely ovate, inconspicuous, 0.7–2.4 mm, membranous.

Flowers

bracteoles 2;

tepals greenish, lanceolate, 3.8–7(–8.5) mm;

inner series slightly shorter, apex sometimes obtuse;

stamens 3–6, filaments (0.7–)1–1.8 mm, anthers 0.3–0.8 mm;

style 0.1–0.2 mm.

tepals reddish to black, narrowly lanceolate to oblong, 2–4.6 × 0.5–1.4 mm;

inner series usually slightly longer than outer, apex obtuse to acuminate;

stamens 3, filaments 0.5–1.4 mm, anthers 1–3.2 mm;

style persistent, 1.4–4 mm, stigma 1.3–4 mm.

Capsules

tan to reddish brown, 3-locular, ellipsoid to narrowly so, slightly truncate, 2.7–4 × 1–1.5 mm, sometimes exceeding inner tepals but usually not outer series.

reddish, 3-locular, globose to ellipsoid or oblong, 2–4.5 × 1–3 mm, nearly equal to sepals.

Seeds

yellowish, widely ellipsoid to ovoid, 0.26–0.49, not tailed.

ovoid, 0.35–0.45 mm.

= 16.

= 27–37, 58–81, 108–115.

Juncus leiospermus

Phenology

Flowering and fruiting spring–early fall.

Flowering and fruiting in spring.

Habitat

Moist soils in meadows, along lakeshores or stream banks, ditches, or roadsides, especially frequent in drawdown areas, usually in open sites and often becoming weedy

Margins of vernal pools

Elevation

0–500 m (0–1600 ft)

Distribution

AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; Greenland; nearly worldwide

[WildflowerSearch map]

[BONAP county map]

[WildflowerSearch map]

[BONAP county map]

Discussion

Nearly worldwide, Juncus bufonius is found essentially throughout North America except north of the Alaskan and Canadian tTaiga. Juncus bufonius is a highly polymorphic complex that is poorly understood systematically. Insufficient evidence exists upon which to base the segregation of the plethora of taxa that have been recognized out of this group in the past.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants from Butte and Calaveras Ccounties, California, tend to be smaller and consistently develop only one flower per culm; they may be recognized as Juncus leiospermus var. ahartii Ertter.

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 22.

Parent taxa

Juncaceae > Juncus > subg. Poiophylli

Juncaceae > Juncus > subg. Graminifolii

Sibling taxa

J. acuminatus, J. acutiflorus, J. acutus, J. alpinoarticulatus, J. anthelatus, J. arcticus, J. articulatus, J. biglumis, J. bolanderi, J. brachycarpus, J. brachycephalus, J. brachyphyllus, J. brevicaudatus, J. bryoides, J. bulbosus, J. caesariensis, J. canadensis, J. capillaris, J. capitatus, J. castaneus, J. chlorocephalus, J. compressus, J. confusus, J. cooperi, J. coriaceus, J. covillei, J. debilis, J. dichotomus, J. diffusissimus, J. drummondii, J. dubius, J. dudleyi, J. effusus, J. elliottii, J. ensifolius, J. falcatus, J. filiformis, J. filipendulus, J. georgianus, J. gerardii, J. greenei, J. gymnocarpus, J. hallii, J. hemiendytus, J. howellii, J. inflexus, J. interior, J. kelloggii, J. leiospermus, J. lesueurii, J. longistylis, J. luciensis, J. macrandrus, J. macrophyllus, J. marginatus, J. maritimus, J. megacephalus, J. mertensianus, J. militaris, J. nevadensis, J. nodatus, J. nodosus, J. occidentalis, J. orthophyllus, J. oxymeris, J. parryi, J. patens, J. pelocarpus, J. pervetus, J. phaeocephalus, J. polycephalus, J. regelii, J. repens, J. roemerianus, J. scirpoides, J. secundus, J. squarrosus, J. stygius, J. subcaudatus, J. subtilis, J. supiniformis, J. tenuis, J. texanus, J. textilis, J. tiehmii, J. torreyi, J. trifidus, J. triformis, J. triglumis, J. trigonocarpus, J. uncialis, J. validus, J. vaseyi, J. xiphioides

J. acuminatus, J. acutiflorus, J. acutus, J. alpinoarticulatus, J. anthelatus, J. arcticus, J. articulatus, J. biglumis, J. bolanderi, J. brachycarpus, J. brachycephalus, J. brachyphyllus, J. brevicaudatus, J. bryoides, J. bufonius, J. bulbosus, J. caesariensis, J. canadensis, J. capillaris, J. capitatus, J. castaneus, J. chlorocephalus, J. compressus, J. confusus, J. cooperi, J. coriaceus, J. covillei, J. debilis, J. dichotomus, J. diffusissimus, J. drummondii, J. dubius, J. dudleyi, J. effusus, J. elliottii, J. ensifolius, J. falcatus, J. filiformis, J. filipendulus, J. georgianus, J. gerardii, J. greenei, J. gymnocarpus, J. hallii, J. hemiendytus, J. howellii, J. inflexus, J. interior, J. kelloggii, J. lesueurii, J. longistylis, J. luciensis, J. macrandrus, J. macrophyllus, J. marginatus, J. maritimus, J. megacephalus, J. mertensianus, J. militaris, J. nevadensis, J. nodatus, J. nodosus, J. occidentalis, J. orthophyllus, J. oxymeris, J. parryi, J. patens, J. pelocarpus, J. pervetus, J. phaeocephalus, J. polycephalus, J. regelii, J. repens, J. roemerianus, J. scirpoides, J. secundus, J. squarrosus, J. stygius, J. subcaudatus, J. subtilis, J. supiniformis, J. tenuis, J. texanus, J. textilis, J. tiehmii, J. torreyi, J. trifidus, J. triformis, J. triglumis, J. trigonocarpus, J. uncialis, J. validus, J. vaseyi, J. xiphioides

Synonyms

J. bufonius var. congestus, J. bufonius var. halophilus, J. bufonius var. hybridus, J. bufonius var. occidentalis, J. bufonius var. ranarius, J. congestus, J. ranarius

J. leiospermus var. ahartii

Name authority

Linnaeus: Sp. Pl. 1: 328. (1753)

F. J. Hermann: Leaflets of Western Botany 5: 113. (1948)

Web links

Local floras: BC, CA, OR, WA
Local Web sites: CalFlora, CalPhotos, Flora NW, Go Botany, IL Wildflowers, KS Wildflowers, LA Plants, MD Biodiversity, MI Flora, MN Wildflowers, MO Plants, PNW Herbaria
WildflowerSearch
iNaturalist (observations)
USDA Plants Database
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Juncus bufonius

Juncus leiospermus

Juncus bufonius

Juncus leiospermus