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jepsonia

foothill jepsonia, foothills jepsonia

Habit Herbs, (acaulescent, heterostylous), not rhizomatous, not stoloniferous; caudex branched or unbranched, ovoid or flat, cormlike, fleshy, without scales.
Caudices

branched, flat.

Flowering stems

(usually appearing in autumn or winter after basal leaves have withered), erect or ascending, leafless, 3–28 cm, stipitate-glandular, slightly viscid.

Leaves

in basal rosette and cauline;

stipules present;

petiole stipitate-glandular;

blade ovate to suborbiculate or orbiculate-reniform, shallowly lobed, base cordate, ultimate margins crenate with callous-apiculate teeth, apex obtuse to rounded, surfaces hairy;

venation palmate.

2–3.

Inflorescences

cymose panicles, (origin of inflorescences not readily resolvable), 2–17-flowered, sometimes flowers solitary, bracteate, (simple or branched distally, glandular-viscid or glabrate).

diffuse, (3–)4–17(–25)-flowered;

bracts scalelike;

peduncle pink or reddish, drying tan or reddish, branched near middle, 3–23 cm.

Flowers

heterostylous;

hypanthium free from ovary except at base, yellow-green to pinkish;

sepals 5, yellow-green to pinkish;

petals 5, white, veins prominently colored, (spatulate to elliptic, glabrous);

nectary tissue not visible;

stamens 10;

filaments subulate, (alternate ones surpassing sepals);

ovary superior, 2-locular, carpels connate to middle;

placentation axile (appearing marginal);

styles 2;

stigmas 2.

hypanthium campanulate, 1.5–3 mm, length to 1.5 times sepals;

sepals pink, 1.3–2 mm, base flattened, glandular-viscid;

petals withering, alternate with sepals, distinct, white with deep pink veins, 3.5–6 mm.

Capsules

folliclelike, 2-beaked.

whitish green or red with red striations, ovoid or ellipsoid, beaks divergent.

Seeds

brownish, irregular in outline with pinched, curved base, reticulate with longitudinal wings or ridges.

light brown.

Pollen

bluish or cream.

x

= 7.

2n

= 14.

Jepsonia

Jepsonia heterandra

Phenology Flowering Sep–Nov.
Habitat Crevices, slate outcrops, rocky slopes
Elevation 50-600 m (200-2000 ft)
Distribution
from FNA
CA; nw Mexico
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Species 3 (3 in the flora).

Jepsonia is characterized by heterostylous flowers that are produced in the autumn after summer drought; some flowers have long styles and short stamens; others have short styles and long stamens. The floral differences associated with heterostyly within populations in this genus are part of an incompatibility mechanism that prevents self-fertilization. Leaves appear after flowering and persist through the winter if moisture is available. Rarely, leaves and flowers are present simultaneously. All three species of Jepsonia also produce a secondary taproot each year. This structure begins to grow after seasonal rains and shrivels before flowering begins. It apparently acts as a contractile root. The three species in this genus are narrow allopatric endemics.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Jepsonia heterandra is relatively uncommon and is apparently restricted to crevices and alluvium of very specific rock types. It has been suggested (R. Ornduff 1969b) that both the mineral content and texture of the substrate influence the distribution of this species. Heterostyly in the genus was first recognized by Eastwood in her description of this species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Caudices unbranched, ovoid to spheric; leaves 1(-3); inflorescences 1-5(-10)-flowered; hypanthia 2+ times sepals; sw California, n Baja California.
J. parryi
1. Caudices branched, ovoid or flat; leaves 2-3; inflorescences (3-)4-17(-25)-flowered; hypanthia ± equaling or to 1.5 times sepals; Sierra Nevada foothills or Channel Islands
→ 2
2. Peduncles pink or reddish, branched near middle; petals withering; pollen bluish or cream; seeds light brown.
J. heterandra
2. Peduncles greenish or olive, branched distally; petals persistent; pollen yellowish; seeds dark brown.
J. malvifolia
Source FNA vol. 8, p. 118. Author: Debra K. Trock. FNA vol. 8, p. 119.
Parent taxa Saxifragaceae Saxifragaceae > Jepsonia
Sibling taxa
J. malvifolia, J. parryi
Subordinate taxa
J. heterandra, J. malvifolia, J. parryi
Synonyms J. parryi var. heterandra
Name authority Small: Bull. Torrey Bot. Club 23: 18, plate 256. 1896 , Eastwood: Bull. Torrey Bot. Club 32: 201. 1905 ,
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