Jarava speciosa |
Poaceae subfam. Pooideae(synonym of Poaceae subfam. pooideae) |
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desert needlegrass |
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Habit | Plants tightly cespitose, not rhizomatous. | Plants annual or perennial; sometimes matlike, sometimes cespitose, sometimes stoloniferous, sometimes rhizomatous. | ||||||||||||||||||||||||||||||||||||
Culms | 30-60 cm, bases orange-brown; nodes 3-6; basal branching intravaginal. |
usually hollow, sometimes solid. |
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Sheaths | mostly glabrous, throats densely ciliate, basal sheaths reddish brown, flat and ribbonlike with age; ligules varying within a plant, lower ligules 0.3-1 mm, densely hairy and ciliate, hairs 0.2-1 mm, often longer than the basal membrane, upper ligules to 2.5 mm, hyaline to scarious, glabrous or hairy, usually less hairy than the lower ligules, sometimes ciliate; blades 10-30 cm long, 0.5-2 mm wide when flat, usually rolled, to 1 mm in diameter, abaxial surfaces glabrous, smooth, adaxial surfaces pilose. |
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Leaves | distichous; sheaths usually open to the base, varying to closed for nearly their full length; auricles present or absent; abaxial ligules absent; adaxial ligules scarious or membranous, sometimes puberulent or scabridulous, usually not ciliate, cilia sometimes shorter than the base; pseudopetioles rarely present; blades usually linear, sometimes broadly so, venation parallel; cross sections non-Kranz, mesophyll nonradiate, adaxial palisade layer absent, fusoid and arm cells usually absent; midribs usually simple; adaxial bulliform cells present; stomates with parallel-sided subsidiary cells; epidermes usually lacking bicellular microhairs, sometimes with unicellular microhairs, papillae usually absent, when present, rarely more than 1 per cell. |
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Panicles | 10-15 cm, dense, frequently partially included in the upper leaf sheaths at maturity; branches ascending. |
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Inflorescences | usually terminal, panicles, spikes, or racemes, usually ebracteate; disarticulation usually below the florets, sometimes below the glumes, at the rachis nodes, or at the inflorescence bases. |
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Spikelets | 16-24 mm. |
usually bisexual, infrequently unisexual or mixed, usually laterally compressed or not compressed, occasionally dorsally compressed, with 1-30 sexual florets, distal floret(s) often reduced, infrequently spikelets with 1-2 reduced or staminate basal florets and a single terminal sexual floret. |
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Glumes | linear-lanceolate, glabrous, tapering from below midlength to the narrowly acute apices; lower glumes 16-24 mm, 1-veined; upper glumes 13-19 mm, 3-5-veined; florets (6)8-10 mm; calluses 0.8-1.6(3) mm, sharp; lemmas densely and evenly hairy, hairs about 0.5 mm, without a pappus; awns 35-45(80) mm, once-geniculate, first segment pilose, hairs 3-8 mm, terminal segment glabrous, smooth; paleas 3.2-5.1 mm, 2/5 – 2/3 (4/5) the length of the lemmas, usually hairy, hairs about 0.5 mm. |
usually 2, upper or lower glumes sometimes absent, rarely both glumes absent; lemmas without uncinate hairs, awned or not, awns single, basal to apical; paleas usually well-developed, sometimes reduced or absent; lodicules 2(3), usually lanceolate and broadly membranous distally, rarely truncate and fleshy, usually not veined or obscurely veined, sometimes distinctly veined, sometimes ciliate; anthers (1, 2)3; ovaries glabrous or sometimes hairy distally, sometimes with an apical appendage; haustorial synergids absent; styles (1)2 (-4), bases close together, sometimes fused. |
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Caryopses | hila linear, elliptic, ovate, or punctate; endosperm usually hard, sometimes soft or liquid, with or without lipids, starch grains compound or simple; embryos less than 1/2 the length of the caryopses; epiblasts usually present; scutellar cleft usually absent; mesocotyl internode usually absent; embryonic leaf margins overlapping, x = 7, 10. |
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2n | = 66, 68, about 74. |
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Jarava speciosa |
Poaceae subfam. Pooideae |
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Distribution | ||||||||||||||||||||||||||||||||||||||
Discussion | Jarava speciosa grows on rocky slopes in canyons of arid and semiarid regions of the southwestern United States and northern Mexico, and in Chile and northern to central Argentina. Several varieties are recognized in South America. It is not clear to which of these varieties, if any, the North American plants belong. The reddish brown leaf bases, differing lower and upper ligules, and the pilose, once-geniculate awns make Jarava speciosa an easy species to recognize in North America. It is also an attractive species, well worth cultivating. It prefers open areas with well-drained soils. The growth of young shoots and flowering is stimulated by fire. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The subfamily Pooideae includes approximately 3300 species, making it the largest subfamily in the Poaceae. It reaches its greatest diversity in cool temperate and boreal regions, extending across the tropics only in high mountains. The circumscription and relationships of tribes within the Pooideae are unsettled (see, for example, Catalan et al. 1997, 2004; Soreng and Davis 1998). In this flora, some previously recognized tribes have been combined with the Poeae. Recognition of some of these as subtribes is well supported; among these is the Hainardieae Greuter (which, at the subtribal level, is called the Parapholiinae Caro). Members of other traditional tribal groupings, such as the Aveneae Dumort., appear to be widely dispersed within the Poeae sensu lato. Further work will probably support the division of the expanded Poeae into additional tribes; there is as yet no clear indication as to what the boundaries of such tribes should be. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 181. | FNA vol. 24, p. 57. | ||||||||||||||||||||||||||||||||||||
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Synonyms | Stipa speciosa, Achnatherum speciosum | |||||||||||||||||||||||||||||||||||||
Name authority | (Trin. & Rupr.) Penail. | Benth. | ||||||||||||||||||||||||||||||||||||
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