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desert needlegrass

rice grass

Habit Plants tightly cespitose, not rhizomatous. Plants perennial; cespitose, sometimes rhizomatous, rhizomes forming knotted bases.
Culms

30-60 cm, bases orange-brown;

nodes 3-6;

basal branching intravaginal.

15-200 cm, not branching at the upper nodes;

basal branching intravaginal or extravaginal;

prophylls not evident, shorter than the leaf sheaths.

Sheaths

mostly glabrous, throats densely ciliate, basal sheaths reddish brown, flat and ribbonlike with age;

ligules varying within a plant, lower ligules 0.3-1 mm, densely hairy and ciliate, hairs 0.2-1 mm, often longer than the basal membrane, upper ligules to 2.5 mm, hyaline to scarious, glabrous or hairy, usually less hairy than the lower ligules, sometimes ciliate;

blades 10-30 cm long, 0.5-2 mm wide when flat, usually rolled, to 1 mm in diameter, abaxial surfaces glabrous, smooth, adaxial surfaces pilose.

Leaves

mostly basal, not overwintering;

sheaths open to the base;

cleistogenes not present;

collars with tufts of hair on either side;

auricles absent;

ligules membranous, truncate or shortest in the center and rounded, edges usually ciliate, hairs at the outer edges often longer than the central membranous portion, ligules of the lower leaves glabrous or hairy, sometimes densely hairy, those of the upper leaves glabrous or sparsely hairy;

blades usually convolute, apices narrowly pointed, flag leaves longer than 10 mm.

Panicles

10-15 cm, dense, frequently partially included in the upper leaf sheaths at maturity;

branches ascending.

Inflorescences

panicles, often partially included in the upper leaf sheath;

branches straight.

Spikelets

16-24 mm.

5.5-24 mm, with 1 floret;

rachillas not prolonged beyond the floret;

disarticulation above the glumes, beneath the floret.

Glumes

linear-lanceolate, glabrous, tapering from below midlength to the narrowly acute apices;

lower glumes 16-24 mm, 1-veined;

upper glumes 13-19 mm, 3-5-veined;

florets (6)8-10 mm;

calluses 0.8-1.6(3) mm, sharp;

lemmas densely and evenly hairy, hairs about 0.5 mm, without a pappus;

awns 35-45(80) mm, once-geniculate, first segment pilose, hairs 3-8 mm, terminal segment glabrous, smooth;

paleas 3.2-5.1 mm, 2/5 – 2/3 (4/5) the length of the lemmas, usually hairy, hairs about 0.5 mm.

unequal, usually longer than the floret, sometimes shorter, hyaline, 0-5-veined;

florets narrowly lanceoloid, terete;

calluses 0.2-1.6(3) mm, acute, less than or equaling the floret diameter, antrorsely strigose distally, hairs white;

lemmas thickly membranous, basal 2/3 scabrous or shortly pubescent, distal 1/3 often bearing a pappus of ascending to strongly divergent 3-8 mm hairs, sometimes glabrous or with appressed hairs shorter than 1 mm, margins not or only slightly overlapping at maturity, apices not fused into a crown, lobes to 0.2 mm, with a single, terminal awn, awn-lemma junction conspicuous;

awns 9-45(80) mm, persistent or deciduous, scabrous, weakly once- or twice-geniculate, first segment scabrous or pilose, terminal segment glabrous or pilose, smooth or scabrous;

paleas 1/3 - 1/2 as long as the lemmas, flat between the veins, membranous to hyaline, glabrous or sparsely pubescent, 2-veined, veins poorly developed, apices rounded to irregular;

lodicules 2-3, the third, if present, reduced;

anthers 3.

Caryopses

fusiform, not ribbed;

hila linear, x = unknown.

2n

= 66, 68, about 74.

Jarava speciosa

Jarava

Distribution
[WildflowerSearch map]
from FNA
CA
[BONAP county map]
Discussion

Jarava speciosa grows on rocky slopes in canyons of arid and semiarid regions of the southwestern United States and northern Mexico, and in Chile and northern to central Argentina. Several varieties are recognized in South America. It is not clear to which of these varieties, if any, the North American plants belong.

The reddish brown leaf bases, differing lower and upper ligules, and the pilose, once-geniculate awns make Jarava speciosa an easy species to recognize in North America. It is also an attractive species, well worth cultivating. It prefers open areas with well-drained soils. The growth of young shoots and flowering is stimulated by fire.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Jarava is a South American genus that used to be included in Stipa. Its limits are currently under study. As treated here, it is a genus of approximately 50 species, all of which are native to South America. It includes three groups that have been recognized as subgenera within Stipa sensu lato: Jarava (Ruiz. & Pav.) Trin. & Rupr., Pappostipa Speg., and Ptilostipa Speg. Jacobs and Everett (1997) recommended including only the approximately 14 species of Stipa subg. Jarava in Jarava. This treatment adopts a somewhat broader interpretation and includes the members of the other two subgenera, pending more detailed study of relationships among the American Stipeae. In this respect, the treatment presented here conforms with the treatment by Pefiailillo (2002); it differs in excluding Achnatberum and Amelichloa.

One species of the Pappostipa group, Jarava speciosa, grows as a disjunct in the southwestern United States, its broadest distribution being in South America. Two other species, /. ichu and /. plumosa, both members of Jarava sensu stricto, have been found as escapes from cultivation in California.

Many species of Jarava have conspicuous hairs on the distal portion of the lemma, termed a pappus, or on the first and/or second segment of the awn. These are an adaptation to wind dispersal. Jarava sensu stricto shows an even stronger adaptation to wind dispersal, usually combining a well-developed pappus with light florets less than 5 mm long and 1 mm wide.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Awns 35-80 cm long, the basal segment pilose; ligules of the basal leaves softly and densely hairy
J. speciosa
1. Awns 9-30 mm long, the basal segment scabridulous or smooth; ligules of the basal leaves glabrous or almost so.
→ 2
2. Glumes clearly exceeding the florets; pappus hairs 3-4 mm long
J. ichu
2. Glumes from shorter than to subequal to the florets; pappus hairs 5-8 mm long
J. plumosa
Source FNA vol. 24, p. 181. FNA vol. 24, p. 178. Author: Mirta O. Arriaga;.
Parent taxa Poaceae > subfam. Pooideae > tribe Stipeae > Jarava Poaceae > subfam. Pooideae > tribe Stipeae
Sibling taxa
J. ichu, J. plumosa
Subordinate taxa
J. ichu, J. plumosa, J. speciosa
Synonyms Stipa speciosa, Achnatherum speciosum
Name authority (Trin. & Rupr.) Penail. Ruiz & Pav.
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