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Peruvian needlegrass

rice grass

Habit Plants densely cespitose, not rhizomatous. Plants perennial; cespitose, sometimes rhizomatous, rhizomes forming knotted bases.
Culms

(15)30-100 cm, bases dull brown, glabrous;

nodes 2-4; branching intravaginal.

15-200 cm, not branching at the upper nodes;

basal branching intravaginal or extravaginal;

prophylls not evident, shorter than the leaf sheaths.

Sheaths

mostly glabrous, scabridulous, basal sheaths dull brown;

ligules 0.3-1 mm, truncate, erose, abaxial surfaces glabrous or almost so, ciliate, hairs longest (to 2 mm) towards the sides of the leaves, at the top of the sheaths;

blades (3)10-40 cm long, 0.5-1 mm wide, all alike, straight, erect, convolute, apices sharp.

Leaves

mostly basal, not overwintering;

sheaths open to the base;

cleistogenes not present;

collars with tufts of hair on either side;

auricles absent;

ligules membranous, truncate or shortest in the center and rounded, edges usually ciliate, hairs at the outer edges often longer than the central membranous portion, ligules of the lower leaves glabrous or hairy, sometimes densely hairy, those of the upper leaves glabrous or sparsely hairy;

blades usually convolute, apices narrowly pointed, flag leaves longer than 10 mm.

Panicles

(3)10-25(30) cm, narrow, cylindrical to lanceoloid, dense, from partially to wholly exserted at anthesis, erect or nodding distally;

branches strongly ascending.

Inflorescences

panicles, often partially included in the upper leaf sheath;

branches straight.

Spikelets

5.5-11 mm.

5.5-24 mm, with 1 floret;

rachillas not prolonged beyond the floret;

disarticulation above the glumes, beneath the floret.

Glumes

subequal, clearly exceeding the florets, linear-lanceolate, tapering to attenuate apices;

lower glumes 5.5-11 mm, 1-3-veined;

upper glumes 5-10.5 mm, 3-veined;

florets 2.3-3 mm, cylindrical to fusiform;

calluses 0.2-0.4 mm, acute to broadly acute, strigose;

lemmas hairy throughout, hairs on the lower portion about 0.15 mm, sparse, appressed, pappus hairs 3-4 mm;

awns 9-15 mm, twice-geniculate, first 2 segments twisted, scabridulous;

paleas 1-1.5 mm, sparsely pubescent, 2-veined, apices rounded;

lodicules 2, 0.6-1 mm;

anthers about 0.8 mm.

unequal, usually longer than the floret, sometimes shorter, hyaline, 0-5-veined;

florets narrowly lanceoloid, terete;

calluses 0.2-1.6(3) mm, acute, less than or equaling the floret diameter, antrorsely strigose distally, hairs white;

lemmas thickly membranous, basal 2/3 scabrous or shortly pubescent, distal 1/3 often bearing a pappus of ascending to strongly divergent 3-8 mm hairs, sometimes glabrous or with appressed hairs shorter than 1 mm, margins not or only slightly overlapping at maturity, apices not fused into a crown, lobes to 0.2 mm, with a single, terminal awn, awn-lemma junction conspicuous;

awns 9-45(80) mm, persistent or deciduous, scabrous, weakly once- or twice-geniculate, first segment scabrous or pilose, terminal segment glabrous or pilose, smooth or scabrous;

paleas 1/3 - 1/2 as long as the lemmas, flat between the veins, membranous to hyaline, glabrous or sparsely pubescent, 2-veined, veins poorly developed, apices rounded to irregular;

lodicules 2-3, the third, if present, reduced;

anthers 3.

Caryopses

1.8-2.2 mm long, 0.6-0.7 mm thick, cylindrical.

fusiform, not ribbed;

hila linear, x = unknown.

Jarava ichu

Jarava

Distribution
from FNA
CA
[BONAP county map]
Discussion

Jarava ichu is native to Mexico, Costa Rica, Venezuela, Colombia, Ecuador, Peru, Bolivia, and Argentina. It is abundant in much of this range. In the Flora region, it is sold as an attractive ornamental. The species could become a problem, because it is self-compatible and produces a large quantity of wind-dispersed seeds. In parts of its native range, J. ichu is highly valued for its ability to prevent soil erosion, and for its use in thatch, mats, and basketry. "Ichu" is a term used to describe any bunchgrass in some parts of South America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Jarava is a South American genus that used to be included in Stipa. Its limits are currently under study. As treated here, it is a genus of approximately 50 species, all of which are native to South America. It includes three groups that have been recognized as subgenera within Stipa sensu lato: Jarava (Ruiz. & Pav.) Trin. & Rupr., Pappostipa Speg., and Ptilostipa Speg. Jacobs and Everett (1997) recommended including only the approximately 14 species of Stipa subg. Jarava in Jarava. This treatment adopts a somewhat broader interpretation and includes the members of the other two subgenera, pending more detailed study of relationships among the American Stipeae. In this respect, the treatment presented here conforms with the treatment by Pefiailillo (2002); it differs in excluding Achnatberum and Amelichloa.

One species of the Pappostipa group, Jarava speciosa, grows as a disjunct in the southwestern United States, its broadest distribution being in South America. Two other species, /. ichu and /. plumosa, both members of Jarava sensu stricto, have been found as escapes from cultivation in California.

Many species of Jarava have conspicuous hairs on the distal portion of the lemma, termed a pappus, or on the first and/or second segment of the awn. These are an adaptation to wind dispersal. Jarava sensu stricto shows an even stronger adaptation to wind dispersal, usually combining a well-developed pappus with light florets less than 5 mm long and 1 mm wide.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Awns 35-80 cm long, the basal segment pilose; ligules of the basal leaves softly and densely hairy
J. speciosa
1. Awns 9-30 mm long, the basal segment scabridulous or smooth; ligules of the basal leaves glabrous or almost so.
→ 2
2. Glumes clearly exceeding the florets; pappus hairs 3-4 mm long
J. ichu
2. Glumes from shorter than to subequal to the florets; pappus hairs 5-8 mm long
J. plumosa
Source FNA vol. 24, p. 179. FNA vol. 24, p. 178. Author: Mirta O. Arriaga;.
Parent taxa Poaceae > subfam. Pooideae > tribe Stipeae > Jarava Poaceae > subfam. Pooideae > tribe Stipeae
Sibling taxa
J. plumosa, J. speciosa
Subordinate taxa
J. ichu, J. plumosa, J. speciosa
Name authority Ruiz & Pav. Ruiz & Pav.
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