Ivesia unguiculata
Rosaceae tribe Potentilleae
Yosemite ivesia, Yosemite mousetail
decumbent to ascending, 1–3.5 dm.
alternate, rarely opposite, pinnately (palmately) compound (simple in Alchemilla, Aphanes, and Chamaerhodos);
stipules persistent (absent in Chamaerhodos), adnate to petiole;
venation pinnate or palmate.
(4–)7–15 cm; sheathing base ± glabrous abaxially;
stipules linear, 3–6 mm;
petiole (0.3–)0.5–4(–5) cm, hairs sparse to abundant, ascending to spreading, 1–2 mm;
leaflets 15–20(–25) per side, loosely overlapping, 3–6 mm, lobes 3–8, linear to oblanceolate, hairs sparse to abundant, spreading, 1–2 mm.
3–6.
(15–)30–100(–200)-flowered, (1–)1.5–4(–8) cm diam., flowers mostly arranged in 1–several loose to tight glomerules of 5–10 flowers.
1–3 mm.
6–9 mm diam.;
epicalyx bractlets linear or narrowly lanceolate to elliptic or narrowly oblong, 1.2–2(–3) mm;
hypanthium shallowly turbinate, 1.5–2.5 × 2–3(–3.5) mm, often nearly as deep as wide;
sepals heavily purple-mottled, (1.5–)2–3(–3.5) mm, acute;
petals white, often tinged with pink, oblanceolate to spatulate or obovate, 3–4 mm;
stamens 10–15, filaments filiform, 0.6–1.1 mm, anthers maroon, 0.3–0.5 mm;
carpels (1–)3–9, styles 1.4–2 mm.
perianth and androecium perigynous;
epicalyx bractlets present, sometimes absent;
hypanthium usually patelliform, cupulate, or campanulate, sometimes turbinate, saucer-shaped, flat-bottomed, or subglobose to ellipsoid or ovoid;
torus flat to conic or turbinate, enlarged (absent or reduced in Alchemilla, Aphanes, and Chamaerhodos);
carpels 1–260, styles basal or lateral to subterminal, distinct;
ovules 1(or 2), basal.
aggregated achenes (achenes in Alchemilla and Aphanes);
torus sometimes fleshy;
styles deciduous or persistent, not elongate.
light brown, 1.2–1.5 mm.
Ivesia unguiculata
Rosaceae tribe Potentilleae
Of conservation concern.
Ivesia unguiculata is found in mid-elevation meadows of the central Sierra Nevada, mostly north of the Kings River. The distinctive deep red to purplish coloration of the inflorescence, and the plant in general, can make patches of this species conspicuous as a smoky purplish haze in meadows. The shape and color of the inflorescences are similar to those of the sympatric Horkelia fusca var. parviflora, suggesting shared pollinators.
The description and illustration by J. D. Hooker (1881), supposedly of Potentilla (Ivesia) unguiculata, were based actually on material grown from seed of I. sericoleuca due to initial confusion of the two species (W. H. Brewer et al. 1876–1880, vol. 1).
The type (Kellogg s.n., CAS) of Potentilla ciliata Greene (not Rafinesque) is unquestionably this species; however, the purported locality (Owens Valley, Inyo County) is dubious and most likely an error in the labeling of the specimen by the collector.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 14–22, species ca. 860 (14 genera, 189 species, including 1 hybrid, in the flora area).
The base chromosome number for Potentilleae is mostly x = 7 (8 in Alchemilla and Aphanes; 14 in Comarum).
Variation in the number of genera recognized in Potentilleae is due to differences in generic delimitation between D. Potter et al. (2007) and the authors of Potentilla and segregates here (see 9. Ivesia and 8. Potentilla for discussion). In the former, Duchesnea, Horkelia, Horkeliella, and Ivesia are included within Potentilla. Likewise, Aphanes is included within Alchemilla by Potter et al. while it is kept distinct here.
Potentilla and its segregates and Fragaria are host to Phragmidium rusts, but not the other genera of the tribe.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Shrubs; leaf lobe margins entire; achenes hirsute. | Dasiphora |
1. Herbs, perennial, sometimes annual or biennial, or subshrubs; leaf lobe margins or apices ± toothed, sometimes entire; achenes glabrous (sometimes ± hairy) | → 2 |
2. Petals 0, sepals 4; achenes 1, enclosed in dry, urceolate or subglobose to ellipsoid or ovoid hypanthia | → 3 |
2. Petals and sepals usually 5; achenes 1–260, usually aggregated (sometimes on elongating tori), usually in (± open) patelliform, cupulate, campanulate, or turbinate hypanthia (not enclosed in dry hypanthium) | → 4 |
3. Herbs perennial; leaves basal, blades reniform to orbiculate, palmately lobed, sometimes palmately compound; stamens 4. | Alchemilla |
3. Herbs annual; leaves cauline, blades cuneate, deeply divided into segments, each lobed; stamen 1(or 2). | Aphanes |
4. Leaves all or mostly basal or proximal (if cauline, deeply pinnatifid), ternate or 2–4-ternate (sometimes simple and coarsely toothed apically in Sibbaldia) | → 5 |
4. Leaves basal or cauline, the latter usually reduced distally, odd-pinnate to palmate, rarely ternate or ± bipinnate | → 9 |
5. Tori becoming red and fleshy in fruit; leaf margins serrate to crenate | → 6 |
5. Tori hemispheric (not enlarged or fleshy) in fruit or absent; leaf margins entire or (2–)3(–5)-toothed apically | → 7 |
6. Leaves ± doubly serrate or crenate; stolons leafy; inflorescences: flowers solitary, axillary at stolon nodes; petals yellow. | Duchesnea |
6. Leaves serrate to crenate; stolons not leafy; inflorescences 1–10-flowered, cymes, axillary from leaf rosettes; petals usually white. | Fragaria |
7. Leaves pinnately compound or simple and deeply pinnatifid, margins entire, stipules absent. | Chamaerhodos |
7. Leaves ternate, margins toothed apically, stipules persistent | → 8 |
8. Petals ± yellow; stamens 5. | Sibbaldia |
8. Petals usually white; stamens 20(–30). | Sibbaldiopsis |
9. Petals deep red to purple, rarely pink, shorter than sepals; tori enlarged and spongy at maturity; horizontal stems sometimes floating, wetland habitats. | Comarum |
9. Petals yellow to white, rarely pink or red (then equal to or longer than sepals); tori not enlarged and spongy at maturity; stems erect to decumbent, not horizontal or floating even if in wetlands | → 10 |
10. Anthers dehiscing by continuous marginal slit (with a single theca); styles sub-basal. | Drymocallis |
10. Anthers dehiscing longitudinally; styles subterminal to lateral | → 11 |
11. Hypanthium patelliform to campanulate or cupulate to turbinate (not flat-bottomed); filaments not forming tube; petals white to yellow, sometimes reddish or pink tinged | → 12 |
11. Hypanthium ± cupulate or bluntly campanulate and flat-bottomed; filaments forming tube; petals usually white, sometimes pink-tinged, rose-veined, or cream | → 13 |
12. Plants not aromatic; leaves ± cordate or reniform to narrowly elliptic in outline, leaflets 3–15(–41); petals oblanceolate or obovate to obcordate to nearly round, rarely elliptic; carpels 3–260. | Potentilla |
12. Plants often aromatic; leaves planar to cylindric, leaflets (3–)7–161; petals linear or narrowly oblanceolate to obovate, sometimes obcordate; carpels 1–20(–40). | Ivesia |
13. Stamens 10; leaflets (3–)5–41. | Horkelia |
13. Stamens 20; leaflets 30–70. | Horkeliella |
CA
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