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Yosemite ivesia, Yosemite mousetail

Bailey's ivesia, Owyhee ivesia

Habit Plants ± grayish, often purple-tinged; glands sparse to abundant. Plants green, ± tufted, often forming hanging clumps, sometimes rosetted.
Stems

decumbent to ascending, 1–3.5 dm.

pendent or prostrate to ascending, 0.5–2(–2.5) dm.

Basal leaves

(4–)7–15 cm; sheathing base ± glabrous abaxially;

stipules linear, 3–6 mm;

petiole (0.3–)0.5–4(–5) cm, hairs sparse to abundant, ascending to spreading, 1–2 mm;

leaflets 15–20(–25) per side, loosely overlapping, 3–6 mm, lobes 3–8, linear to oblanceolate, hairs sparse to abundant, spreading, 1–2 mm.

planar, 3–12(–18) cm; sheathing base not or sparsely strigose abaxially;

petiole 1–8 cm;

lateral leaflets 2–6(–10) per side, separate to slightly overlapping distally, ovate or obovate to flabellate, 4–15(–25) mm, incised 1/4–3/4 to base into 3–11(–15) ovate to oblanceolate teeth or lobes, apex not setose, surfaces ± sparsely short-pilose or hirsute, ± glandular;

terminal leaflets ± distinct.

Cauline leaves

3–6.

1–2;

blade reduced.

Inflorescences

(15–)30–100(–200)-flowered, (1–)1.5–4(–8) cm diam., flowers mostly arranged in 1–several loose to tight glomerules of 5–10 flowers.

(1–)5–40-flowered, open, (1–)1.5–8(–10) cm diam.

Pedicels

1–3 mm.

2–15(–30) mm.

Flowers

6–9 mm diam.;

epicalyx bractlets linear or narrowly lanceolate to elliptic or narrowly oblong, 1.2–2(–3) mm;

hypanthium shallowly turbinate, 1.5–2.5 × 2–3(–3.5) mm, often nearly as deep as wide;

sepals heavily purple-mottled, (1.5–)2–3(–3.5) mm, acute;

petals white, often tinged with pink, oblanceolate to spatulate or obovate, 3–4 mm;

stamens 10–15, filaments filiform, 0.6–1.1 mm, anthers maroon, 0.3–0.5 mm;

carpels (1–)3–9, styles 1.4–2 mm.

4–10 mm diam.;

epicalyx bractlets 5, narrowly lanceolate to ovate, 0.8–2.5 mm;

hypanthium patelliform, 0.5–2 × 2–4 mm;

sepals (1.2–)1.5–4 mm, acute;

petals white or pale yellow, oblanceolate to narrowly spatulate, 1.5–2.5 mm;

stamens 5, filaments 0.5–1.1 mm, anthers yellow, sometimes with reddish margins, oblong, 0.4–0.7 mm;

carpels (1–)3–8, styles 0.9–1.8 mm.

Achenes

light brown, 1.2–1.5 mm.

greenish white to light tan, 1.5–2 mm, smooth or rugose, ± carunculate.

Ivesia unguiculata

Ivesia baileyi

Phenology Flowering summer.
Habitat Moist meadows and slopes, in montane conifer woodlands
Elevation 1500–2500 m (4900–8200 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA; ID; NV; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Ivesia unguiculata is found in mid-elevation meadows of the central Sierra Nevada, mostly north of the Kings River. The distinctive deep red to purplish coloration of the inflorescence, and the plant in general, can make patches of this species conspicuous as a smoky purplish haze in meadows. The shape and color of the inflorescences are similar to those of the sympatric Horkelia fusca var. parviflora, suggesting shared pollinators.

The description and illustration by J. D. Hooker (1881), supposedly of Potentilla (Ivesia) unguiculata, were based actually on material grown from seed of I. sericoleuca due to initial confusion of the two species (W. H. Brewer et al. 1876–1880, vol. 1).

The type (Kellogg s.n., CAS) of Potentilla ciliata Greene (not Rafinesque) is unquestionably this species; however, the purported locality (Owens Valley, Inyo County) is dubious and most likely an error in the labeling of the specimen by the collector.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

The two varieties accepted here were not recognized by D. D. Keck (1938), who instead treated Ivesia setosa as a variety of I. baileyi. Field investigations confirm that the three entities are reasonably distinct morphologically, with intergradation where their otherwise distinct ranges overlap in central Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals pale yellow; hypanthium interior yellow to golden; epicalyx bractlets ± 3/4 to as long as sepals; achenes smooth.
var. baileyi
1. Petals white; hypanthium interior pale green or cream to maroon; epicalyx bractlets usually less than 1/2 as long as sepals; achenes rugose.
var. beneolens
Source FNA vol. 9, p. 241. FNA vol. 9, p. 226.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Unguiculatae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Setosae
Sibling taxa
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. utahensis, I. webberi
I. aperta, I. argyrocoma, I. arizonica, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
Subordinate taxa
I. baileyi var. baileyi, I. baileyi var. beneolens
Synonyms Potentilla unguiculata Potentilla baileyi
Name authority A. Gray: Proc. Amer. Acad. Arts 7: 339. (1868) S. Watson: Botany (Fortieth Parallel), 90. (1871)
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