FNA: Ivesia setosa vs. Ivesia unguiculata

	Ivesia setosa	Ivesia unguiculata
	bristly ivesia	Yosemite ivesia, Yosemite mousetail
Habit	Plants green, tufted to ± densely matted.	Plants ± grayish, often purple-tinged; glands sparse to abundant.
Stems	± ascending to nearly erect, 0.7–2.5(–2.8) dm.	decumbent to ascending, 1–3.5 dm.
Basal leaves	weakly planar to loosely cylindric, 5–9(–12) cm; sheathing base ± strigose abaxially; petiole 1–8 cm; lateral leaflets 5–10 per side, separate to overlapping distally, ± flabellate, 2–8 mm, incised ± 3/4 to base, sometimes nearly to base, into (3–)7–11 ovate teeth to narrowly obovate lobes, apex usually ± setose, surfaces ± sparsely hirsute, conspicuously glandular; terminal leaflets ± indistinct.	(4–)7–15 cm; sheathing base ± glabrous abaxially; stipules linear, 3–6 mm; petiole (0.3–)0.5–4(–5) cm, hairs sparse to abundant, ascending to spreading, 1–2 mm; leaflets 15–20(–25) per side, loosely overlapping, 3–6 mm, lobes 3–8, linear to oblanceolate, hairs sparse to abundant, spreading, 1–2 mm.
Cauline leaves	(0–)1; blade vestigial.	3–6.
Inflorescences	(1–)5–15(–30)-flowered, ± open, 1–6(–12) cm diam.	(15–)30–100(–200)-flowered, (1–)1.5–4(–8) cm diam., flowers mostly arranged in 1–several loose to tight glomerules of 5–10 flowers.
Pedicels	5–15(–20) mm.	1–3 mm.
Flowers	7–10 mm diam.; epicalyx bractlets 5, lanceolate, 1.3–2.5(–3) mm; hypanthium patelliform, (1–)1.5–2 × 2–3.5(–4) mm; sepals (1.5–)2–3.5 mm, ± acute; petals yellow, oblanceolate to narrowly spatulate, 1.5–2.5 mm; stamens 5, filaments 0.8–1.5 mm, anthers yellow, oblong, 0.4–0.7 mm; carpels 2–8, styles 1.2–2 mm.	6–9 mm diam.; epicalyx bractlets linear or narrowly lanceolate to elliptic or narrowly oblong, 1.2–2(–3) mm; hypanthium shallowly turbinate, 1.5–2.5 × 2–3(–3.5) mm, often nearly as deep as wide; sepals heavily purple-mottled, (1.5–)2–3(–3.5) mm, acute; petals white, often tinged with pink, oblanceolate to spatulate or obovate, 3–4 mm; stamens 10–15, filaments filiform, 0.6–1.1 mm, anthers maroon, 0.3–0.5 mm; carpels (1–)3–9, styles 1.4–2 mm.
Achenes	greenish white to light tan, 1.7–2 mm, smooth, ± carunculate.	light brown, 1.2–1.5 mm.

	Ivesia setosa	Ivesia unguiculata
Phenology	Flowering summer.	Flowering summer.
Habitat	Dry, rocky talus slopes, boulders and outcrops, most often but not always of calcareous origin, occasionally away from immediate outcrops, sagebrush communities, conifer woodlands	Moist meadows and slopes, in montane conifer woodlands
Elevation	1800–2600(–3100) m (5900–8500(–10200) ft)	1500–2500 m (4900–8200 ft)
Distribution	NV; UT	CA [BONAP county map]
Discussion	Ivesia setosa occurs mostly to the east and south of I. baileyi, from southeastern Humboldt and Churchill counties to Elko, White Pine, and northern Nye counties, Nevada, and in the Deep Creek Range of western Utah. It represents a transition between the planar-leaved, chasmophytic members of sect. Setosae and the cylindric-leaved, matted species of flatter sites. Stems of I. setosa are more generally erect than in other species in the section, and the usually calcareous substrate is noteworthy. The deeply incised (usually not quite to base) leaflets are somewhat intermediate between the toothed leaflets of I. baileyi and the leaflets of I. shockleyi that are incised to the base into separate lobes. The individual leaflets of both I. baileyi and I. setosa are more or less flat and distichously paired; in I. shockleyi, groups of leaflet lobes are folded over onto each other, giving a verticillate appearance to the leaflet arrangement. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Of conservation concern. Ivesia unguiculata is found in mid-elevation meadows of the central Sierra Nevada, mostly north of the Kings River. The distinctive deep red to purplish coloration of the inflorescence, and the plant in general, can make patches of this species conspicuous as a smoky purplish haze in meadows. The shape and color of the inflorescences are similar to those of the sympatric Horkelia fusca var. parviflora, suggesting shared pollinators. The description and illustration by J. D. Hooker (1881), supposedly of Potentilla (Ivesia) unguiculata, were based actually on material grown from seed of I. sericoleuca due to initial confusion of the two species (W. H. Brewer et al. 1876–1880, vol. 1). The type (Kellogg s.n., CAS) of Potentilla ciliata Greene (not Rafinesque) is unquestionably this species; however, the purported locality (Owens Valley, Inyo County) is dubious and most likely an error in the labeling of the specimen by the collector. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 9, p. 227.	FNA vol. 9, p. 241.
Parent taxa	Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Setosae	Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Unguiculatae
Sibling taxa	I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi	I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. utahensis, I. webberi
Synonyms	I. baileyi var. setosa, Potentilla baileyi var. setosa	Potentilla unguiculata
Name authority	(S. Watson) Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 290. (1908)	A. Gray: Proc. Amer. Acad. Arts 7: 339. (1868)
Web links	Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Ivesia unguiculata

bristly ivesia

Yosemite ivesia, Yosemite mousetail

Habit

Plants green, tufted to ± densely matted.

Plants ± grayish, often purple-tinged; glands sparse to abundant.

Stems

± ascending to nearly erect, 0.7–2.5(–2.8) dm.

decumbent to ascending, 1–3.5 dm.

Basal leaves

weakly planar to loosely cylindric, 5–9(–12) cm; sheathing base ± strigose abaxially;

petiole 1–8 cm;

lateral leaflets 5–10 per side, separate to overlapping distally, ± flabellate, 2–8 mm, incised ± 3/4 to base, sometimes nearly to base, into (3–)7–11 ovate teeth to narrowly obovate lobes, apex usually ± setose, surfaces ± sparsely hirsute, conspicuously glandular;

terminal leaflets ± indistinct.

(4–)7–15 cm; sheathing base ± glabrous abaxially;

stipules linear, 3–6 mm;

petiole (0.3–)0.5–4(–5) cm, hairs sparse to abundant, ascending to spreading, 1–2 mm;

leaflets 15–20(–25) per side, loosely overlapping, 3–6 mm, lobes 3–8, linear to oblanceolate, hairs sparse to abundant, spreading, 1–2 mm.

Cauline leaves

(0–)1;

blade vestigial.

3–6.

Inflorescences

(1–)5–15(–30)-flowered, ± open, 1–6(–12) cm diam.

(15–)30–100(–200)-flowered, (1–)1.5–4(–8) cm diam., flowers mostly arranged in 1–several loose to tight glomerules of 5–10 flowers.

Pedicels

5–15(–20) mm.

1–3 mm.

Flowers

7–10 mm diam.;

epicalyx bractlets 5, lanceolate, 1.3–2.5(–3) mm;

hypanthium patelliform, (1–)1.5–2 × 2–3.5(–4) mm;

sepals (1.5–)2–3.5 mm, ± acute;

petals yellow, oblanceolate to narrowly spatulate, 1.5–2.5 mm;

stamens 5, filaments 0.8–1.5 mm, anthers yellow, oblong, 0.4–0.7 mm;

carpels 2–8, styles 1.2–2 mm.

6–9 mm diam.;

epicalyx bractlets linear or narrowly lanceolate to elliptic or narrowly oblong, 1.2–2(–3) mm;

hypanthium shallowly turbinate, 1.5–2.5 × 2–3(–3.5) mm, often nearly as deep as wide;

sepals heavily purple-mottled, (1.5–)2–3(–3.5) mm, acute;

petals white, often tinged with pink, oblanceolate to spatulate or obovate, 3–4 mm;

stamens 10–15, filaments filiform, 0.6–1.1 mm, anthers maroon, 0.3–0.5 mm;

carpels (1–)3–9, styles 1.4–2 mm.

Achenes

greenish white to light tan, 1.7–2 mm, smooth, ± carunculate.

light brown, 1.2–1.5 mm.

Ivesia setosa

Ivesia unguiculata

Phenology

Flowering summer.

Habitat

Dry, rocky talus slopes, boulders and outcrops, most often but not always of calcareous origin, occasionally away from immediate outcrops, sagebrush communities, conifer woodlands

Moist meadows and slopes, in montane conifer woodlands

Elevation

1800–2600(–3100) m (5900–8500(–10200) ft)

1500–2500 m (4900–8200 ft)

Distribution

NV; UT

[BONAP county map]

Discussion

Ivesia setosa occurs mostly to the east and south of I. baileyi, from southeastern Humboldt and Churchill counties to Elko, White Pine, and northern Nye counties, Nevada, and in the Deep Creek Range of western Utah. It represents a transition between the planar-leaved, chasmophytic members of sect. Setosae and the cylindric-leaved, matted species of flatter sites. Stems of I. setosa are more generally erect than in other species in the section, and the usually calcareous substrate is noteworthy. The deeply incised (usually not quite to base) leaflets are somewhat intermediate between the toothed leaflets of I. baileyi and the leaflets of I. shockleyi that are incised to the base into separate lobes. The individual leaflets of both I. baileyi and I. setosa are more or less flat and distichously paired; in I. shockleyi, groups of leaflet lobes are folded over onto each other, giving a verticillate appearance to the leaflet arrangement.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Ivesia unguiculata is found in mid-elevation meadows of the central Sierra Nevada, mostly north of the Kings River. The distinctive deep red to purplish coloration of the inflorescence, and the plant in general, can make patches of this species conspicuous as a smoky purplish haze in meadows. The shape and color of the inflorescences are similar to those of the sympatric Horkelia fusca var. parviflora, suggesting shared pollinators.

The description and illustration by J. D. Hooker (1881), supposedly of Potentilla (Ivesia) unguiculata, were based actually on material grown from seed of I. sericoleuca due to initial confusion of the two species (W. H. Brewer et al. 1876–1880, vol. 1).

The type (Kellogg s.n., CAS) of Potentilla ciliata Greene (not Rafinesque) is unquestionably this species; however, the purported locality (Owens Valley, Inyo County) is dubious and most likely an error in the labeling of the specimen by the collector.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 9, p. 227.

FNA vol. 9, p. 241.

Parent taxa

Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Setosae

Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Unguiculatae

Sibling taxa

I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi

I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. utahensis, I. webberi

Synonyms

I. baileyi var. setosa, Potentilla baileyi var. setosa

Potentilla unguiculata

Name authority

(S. Watson) Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 290. (1908)

A. Gray: Proc. Amer. Acad. Arts 7: 339. (1868)

Web links

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Ivesia setosa

Ivesia unguiculata

Ivesia setosa

Ivesia unguiculata