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bristly ivesia

Coconino ivesia, manyleaf mousetail, red comarella

Habit Plants green, tufted to ± densely matted. Plants green.
Stems

± ascending to nearly erect, 0.7–2.5(–2.8) dm.

ascending to erect, 2–6 dm.

Basal leaves

weakly planar to loosely cylindric, 5–9(–12) cm; sheathing base ± strigose abaxially;

petiole 1–8 cm;

lateral leaflets 5–10 per side, separate to overlapping distally, ± flabellate, 2–8 mm, incised ± 3/4 to base, sometimes nearly to base, into (3–)7–11 ovate teeth to narrowly obovate lobes, apex usually ± setose, surfaces ± sparsely hirsute, conspicuously glandular;

terminal leaflets ± indistinct.

6–25(–30) cm; sheathing base minutely glandular abaxially;

petiole 0.5–8(–10) cm;

leaflets 12–30 per side, obovate to broadly cuneate, 5–12(–15) mm, incised 1/3–1/2 to base into 3–5 oblong to ovate teeth, sometimes entire, sparsely short-strigose to -villous, sometimes glabrate.

Cauline leaves

(0–)1;

blade vestigial.

Inflorescences

(1–)5–15(–30)-flowered, ± open, 1–6(–12) cm diam.

(5–)8–60-flowered, 2–12 cm diam.

Pedicels

5–15(–20) mm.

5–30(–35) mm.

Flowers

7–10 mm diam.;

epicalyx bractlets 5, lanceolate, 1.3–2.5(–3) mm;

hypanthium patelliform, (1–)1.5–2 × 2–3.5(–4) mm;

sepals (1.5–)2–3.5 mm, ± acute;

petals yellow, oblanceolate to narrowly spatulate, 1.5–2.5 mm;

stamens 5, filaments 0.8–1.5 mm, anthers yellow, oblong, 0.4–0.7 mm;

carpels 2–8, styles 1.2–2 mm.

9–12 mm diam.;

epicalyx bractlets narrowly lanceolate to lanceolate, 1–2 mm;

hypanthium interior brick- to blood-red or golden, 0.5–1.5 × 3–5 mm;

sepals 3–4.5 mm, base brick- to blood-red adaxially, apex acute to acuminate;

petals brick- to blood-red, linear to oblanceolate or narrowly elliptic, 2–3.5 mm;

stamens 5, filaments 1.3–2 mm, anthers maroon, oblong, 0.6–0.9 mm;

carpels 1–5, styles 2–3 mm.

Achenes

greenish white to light tan, 1.7–2 mm, smooth, ± carunculate.

brown, 2–2.5 mm.

2n

= 28.

Ivesia setosa

Ivesia multifoliolata

Phenology Flowering summer. Flowering summer.
Habitat Dry, rocky talus slopes, boulders and outcrops, most often but not always of calcareous origin, occasionally away from immediate outcrops, sagebrush communities, conifer woodlands Dry flats and slopes, on gravelly volcanic or sandstone soil, often along seasonal stream courses, in oak and conifer woodlands
Elevation 1800–2600(–3100) m (5900–8500(–10200) ft) 1800–2400 m (5900–7900 ft)
Distribution
from FNA
NV; UT
from FNA
AZ
[BONAP county map]
Discussion

Ivesia setosa occurs mostly to the east and south of I. baileyi, from southeastern Humboldt and Churchill counties to Elko, White Pine, and northern Nye counties, Nevada, and in the Deep Creek Range of western Utah. It represents a transition between the planar-leaved, chasmophytic members of sect. Setosae and the cylindric-leaved, matted species of flatter sites. Stems of I. setosa are more generally erect than in other species in the section, and the usually calcareous substrate is noteworthy. The deeply incised (usually not quite to base) leaflets are somewhat intermediate between the toothed leaflets of I. baileyi and the leaflets of I. shockleyi that are incised to the base into separate lobes. The individual leaflets of both I. baileyi and I. setosa are more or less flat and distichously paired; in I. shockleyi, groups of leaflet lobes are folded over onto each other, giving a verticillate appearance to the leaflet arrangement.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Ivesia multifoliolata is found between the Grand Canyon and Mogollon Rim in northern Arizona. It is both the only species of the genus with red petals and the only one that overlaps the range of the red-petaled Potentilla sect. Rubrae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 227. FNA vol. 9, p. 244.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Setosae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Comarella
Sibling taxa
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
Synonyms I. baileyi var. setosa, Potentilla baileyi var. setosa Horkelia multifoliolata, Comarella multifoliolata, Potentilla multifoliolata
Name authority (S. Watson) Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 290. (1908) (Torrey) D. D. Keck: Lloydia 1: 125. (1939)
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