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grimy ivesia, grimy mousetail

field ivesia, field mousetail, Kaweah ivesia

Habit Plants grayish, ± matted. Plants green to grayish; glands sparse.
Stems

± prostrate, (0.1–)0.2–1.5(–2) dm.

decumbent to ascending, 1–3.5 dm.

Basal leaves

tightly cylindric, 2–10 cm; sheathing base densely hairy abaxially;

petiole 0.5–4(–10) cm;

lateral leaflets 4–10(–15) per side, overlapping at least distally, ± flabellate, 0.5–3(–4) mm, incised to base or nearly so into (0–)2–4(–9) elliptic to obovate or orbiculate lobes, apex usually not setose, surfaces densely hirsute, cryptically glandular;

terminal leaflets indistinct.

(3–)5–15(–18) cm; sheathing base glabrous or sparsely strigose abaxially;

stipules ± lanceolate, 2.5–5 mm;

petiole (0.3–)0.5–6(–8) cm, hairs sparse to abundant, appressed-ascending, 1–2 mm;

leaflets 15–20 per side, loosely overlapping, 2–10 mm, lobes 2–5, oblanceolate, hairs sparse to abundant, spreading to ascending, 1–2 mm.

Cauline leaves

(0–)1;

blade vestigial.

3–4.

Inflorescences

5–60(–100)-flowered, congested, 0.5–5(–7) cm diam.

5–20(–40)-flowered, (1–)1.5–2.5(–4) cm diam., flowers arranged in 1–few ± tight glomerules of 10–15 flowers.

Pedicels

1.5–4(–8) mm.

1–4 mm.

Flowers

4–8 mm diam.;

epicalyx bractlets 5, linear to elliptic or narrowly ovate, 0.8–1.6(–2.1) mm;

hypanthium patelliform to shallowly cupulate, 1–1.5 × (1.5–)2–3(–3.5) mm;

sepals 1.4–2.5(–2.8) mm, acute;

petals white to pale yellowish, linear to narrowly oblanceolate, 1–1.5 mm;

stamens 5, filaments 0.6–1.8 mm, anthers maroon or yellow with maroon margins, oblong, 0.3–0.6 mm;

carpels 1–2(–4), styles 1–1.5 mm.

7–10 mm diam.;

epicalyx bractlets linear to oblong, 1–2(–2.5) mm;

hypanthium campanulate, 1–2 × 2.5–4 mm, ± 1/2 as deep as wide;

sepals green, 2–3(–3.5) mm, acute;

petals 4(–5), light yellow, oblanceolate to spatulate or narrowly obovate, 3–4 mm;

stamens 12–16 (4-merous flowers) or 16–20 (5-merous flowers), filaments filiform, 0.6–1.1 mm, anthers yellowish, 0.3–0.5 mm;

carpels 4–20, styles 1.4–2 mm.

Achenes

brown, 1.2–1.6(–2) mm, smooth, prominently carunculate.

light brown, 1–1.5 mm.

Ivesia rhypara

Ivesia campestris

Phenology Flowering summer.
Habitat Moist meadows and slopes, in montane to subalpine conifer woodlands
Elevation 2200–3400 m (7200–11200 ft)
Distribution
from FNA
NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Ivesia rhypara forms densely hairy mats or mounds on isolated edaphic sites in the northern Great Basin. Only one of the widely disjunct population clusters is recognized as a distinct variety; the others have general differences in gestalt, but no significant distinctions have been determined (B. Ertter 1989). Isozyme analysis likewise does not show significant differences (T. Kaye et al., unpubl., BLM and Oregon Dept. of Agriculture, 1991). Reproductive studies by D. Wiens (pers. comm.) show low seed set (ca. 5% in var. rhypara, somewhat higher in var. shellyi), possibly indicating a deleterious genetic load.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Ivesia campestris is found in the southern Sierra Nevada south of the Kings River, where it replaces I. unguiculata and often occurs at higher elevations.

Ivesia campestris is the only member of the genus that commonly has 4-merous flowers. It is sometimes misidentified as I. unguiculata, especially when the pale yellow petals have faded to creamy white; the glomerules of the inflorescences tend to be yellowish green to green rather than purplish. Also, the anthers of I. campestris are yellowish, rather than maroon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants (4–)8–30 cm diam.; inflorescences (1–)2–5(–7) cm diam.; scarcely or cryptically petrophytic in bedrock overlain by rubble and soil.
var. rhypara
1. Plants 2–10(–15) cm diam.; inflorescences 0.5–2(–5) cm diam.; overtly petrophytic in boulders and outcrops.
var. shellyi
Source FNA vol. 9, p. 228. FNA vol. 9, p. 242.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Setosae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Unguiculatae
Sibling taxa
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
Subordinate taxa
I. rhypara var. rhypara, I. rhypara var. shellyi
Synonyms Potentilla utahensis var. campestris, P. campestris
Name authority Ertter & Reveal: Madroño 24: 224, fig. 1. (1977) (M. E. Jones) Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 285. (1908)
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