Ivesia paniculata
Ivesia sect. Setosae
Ash Creek ivesia, Ash Creek mousetail
± prostrate, 0.4–1.5(–2) dm.
(0.1–)0.2–2.5(–3) dm.
tightly cylindric, (1.5–)2–5(–7) cm; sheathing base densely hairy abaxially;
petiole 0.5–4 cm;
lateral leaflets (5–)8–15 per side, overlapping at least distally, ± flabellate, 0.5–2 mm, incised to base or nearly so into (0–)3–8(–15) elliptic to narrowly obovate lobes, apex not or obscurely setose, surfaces densely hirsute, cryptically glandular;
terminal leaflets indistinct.
planar or loosely to tightly cylindric;
stipules present;
leaflets separate to overlapping, usually individually distinguishable, toothed 1/4–3/4 or lobed to base, rarely entire, sparsely to densely hairy;
terminal leaflets distinct or indistinct.
(0–)1;
blade reduced.
(0–)1–2(–4), not paired;
blade well developed to vestigial.
20–200-flowered, congested, (1–)1.5–6(–10) cm diam.
open to congested, flowers arranged ± individually (or ± glomerulate in I. paniculata and I. rhypara).
1.5–6 mm.
usually becoming ± curved, often sigmoid.
4–6 mm diam.;
epicalyx bractlets 5, lanceolate to elliptic, 0.6–1.5(–2) mm;
hypanthium shallowly cupulate, 1 × 2–3 mm;
sepals (1–)1.5–2.5(–3) mm, acute;
petals white to pale yellowish, linear, 1 mm;
stamens 5, filaments 0.3–1 mm, anthers yellow with maroon margins, ovate, 0.3–0.5 mm;
carpels 1–2(–3), styles 0.7–1.8 mm.
hypanthium mostly patelliform, sometimes shallowly cupulate or (in I. arizonica) campanulate or turbinate;
petals not medially reflexed, yellow to white, not clawed, apex obtuse to rounded;
stamens 5–35(–40), anthers longer or shorter than wide, laterally dehiscent;
carpels 1–20(–40).
brown, 0.8–1.5 mm, smooth, prominently carunculate.
vertical, usually rugose, sometimes smooth, ± carunculate (except I. cryptocaulis).
Ivesia paniculata
Ivesia sect. Setosae
Of conservation concern.
Ivesia paniculata is known only from the Ash Valley area of Lassen County. The distinctions between I. paniculata and I. rhypara are perhaps on the same scale as variation among disjunct population clusters of I. rhypara, but no taxonomic adjustments are proposed at this time.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Species 11 (11 in the flora).
The planar-leaved, chasmophytic members of sect. Setosae found in and around the mountains centered on the Mojave Desert are hypothesized to represent the ancestral radiation of Ivesia, on the basis of morphology and distribution. A secondary radiation northward into the Great Basin is characterized by a complete sequence in size reduction, dissection, and increased number of leaflets, paralleled by a reduction in numbers of floral parts. Ivesia saxosa retains the most pleisiomorphies and could readily be accommodated within Potentilla in the strict sense, as has commonly been done. Gross morphology of I. saxosa is nearly identical to that of I. arizonica, which has been treated as a distinct genus, Purpusia, on the basis of its lack of an epicalyx, deep hypanthium, and stipelike torus. The narrow endemic I. patellifera is intermediate between these two species, and an identical habit is shared by I. baileyi and I. jaegeri, which have more dissected leaflets and more reduced flowers. All of these species are now placed in the same genus, either as Ivesia (B. Ertter 1989) or as Potentilla (J. T. Howell 1945).
Most species in sect. Setosae are more or less petrophytic (in contrast to other sections), often growing in crevices of vertical cliffs. The protection offered by these habitats often allows extended blooming periods and resultant indeterminate inflorescences that produce flowers as long as conditions are suitable, including at the height of summer when relatively few other desert species are in bloom. Stems can initially radiate in almost all directions relative to the substrate (vertical in chasmophytes, horizontal in others), but they generally succumb to gravity with increased number of flowers and become either pendent or prostrate. In particularly favorable years, blooming season and flower number can exceed the ranges given here. The small but often numerous flowers are visited by a wide diversity of potential pollinators, including ants, which are attracted to the glistening, nectariferous hypanthial disc. Most species of sect. Setosae have carunculate seeds, which may indicate that ants play a role in seed dispersal, a particularly useful adaptation for chasmophytic plants.
Since Ivesia longibracteata (sect. Ivesia) is sometimes identified as a member of sect. Setosae, it is included herein and keys out in the sixth couplet.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Basal leaves planar; lateral leaflets (2–)5–20(–25) mm, incised 1/4–3/4 to base into teeth or lobes | → 2 |
1. Basal leaves ± cylindric to weakly planar; lateral leaflets 0.5–8 mm, incised to base or nearly so into separate lobes, sometimes entire | → 6 |
2. Stamens 15–35(–40). | I. saxosa |
2. Stamens 5–10 | → 3 |
3. Epicalyx bractlets 0(–3); Mojave Desert and n Arizona | → 4 |
3. Epicalyx bractlets 5; n Great Basin and adjacent mountains | → 5 |
4. Hypanthia patelliform, 0.5(–1) mm deep; Kingston Range, California. | I. patellifera |
4. Hypanthia turbinate or campanulate, 1.5–3(–5) mm deep; se California, s Nevada, and n Arizona. | I. arizonica |
5. Lateral leaflets incised 1/4–3/4 to base, lobes not setose apically; basal leaves: sheathing bases not or sparsely strigose abaxially; volcanic substrates. | I. baileyi |
5. Lateral leaflets incised ± 3/4 to base, sometimes nearly to base, lobes usually ± setose apically; basal leaves: sheathing bases ± strigose abaxially; usually calcareous substrates. | I. setosa |
6. Epicalyx bractlets longer than sepals; inflorescences loosely subcapitate; pedicels remaining straight (Castle Crags, n California). | I. longibracteata |
6. Epicalyx bractlets shorter than (or equal to) sepals; inflorescences open to congested, not subcapitate; pedicels usually ± curved in fruit | → 7 |
7. Stamens 20 | → 8 |
7. Stamens 5 | → 9 |
8. Petals yellow, narrowly oblanceolate; lateral leaflet lobes 3–6; Spring Mountains, Nevada, and Clark Mountain, California. | I. jaegeri |
8. Petals white, obovate; lateral leaflets lobes (0–)2–4; San Jacinto Mountains, California. | I. callida |
9. Plants grayish; lateral leaflet surfaces densely hirsute, cryptically glandular; inflorescences congested, 5–200-flowered; petals linear to narrowly oblanceolate, 1–1.5 mm, white to pale yellowish; 1300–1900 m | → 10 |
9. Plants green to grayish green; lateral leaflet surfaces ± sparsely hirsute or loosely long-strigose, evidently glandular; inflorescences ± open to ± congested, 1–20(–30)-flowered; petals oblanceolate to spatulate or narrowly obovate, (1–)1.5–3.2 mm, ± yellow; 1700–4000 m | → 11 |
10. Leaflet lobes (0–)2–4(–9), elliptic to obovate or orbiculate, 0.5–3(–4) mm; n Nevada, se Oregon. | I. rhypara |
10. Leaflet lobes (0–)3–8(–15), elliptic to narrowly obovate, 0.5–2 mm; ne California. | I. paniculata |
11. Plants diffusely matted; lateral leaflet surfaces loosely long-strigose; Spring Mountains, s Nevada. | I. cryptocaulis |
11. Plants tufted to ± densely matted; lateral leaflet surfaces ± sparsely hirsute; Great Basin and n Sierra Nevada | → 12 |
12. Lateral leaflets incised ± 3/4 to base, sometimes nearly to base, into ovate teeth to narrowly obovate lobes; usually calcareous substrates. | I. setosa |
12. Lateral leaflets incised to base or nearly so into oblanceolate to obovate or elliptic lobes; various substrates, seldom calcareous. | I. shockleyi |
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