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Ash Creek ivesia, Ash Creek mousetail

Coconino ivesia, manyleaf mousetail, red comarella

Habit Plants grayish, ± matted. Plants green.
Stems

± prostrate, 0.4–1.5(–2) dm.

ascending to erect, 2–6 dm.

Basal leaves

tightly cylindric, (1.5–)2–5(–7) cm; sheathing base densely hairy abaxially;

petiole 0.5–4 cm;

lateral leaflets (5–)8–15 per side, overlapping at least distally, ± flabellate, 0.5–2 mm, incised to base or nearly so into (0–)3–8(–15) elliptic to narrowly obovate lobes, apex not or obscurely setose, surfaces densely hirsute, cryptically glandular;

terminal leaflets indistinct.

6–25(–30) cm; sheathing base minutely glandular abaxially;

petiole 0.5–8(–10) cm;

leaflets 12–30 per side, obovate to broadly cuneate, 5–12(–15) mm, incised 1/3–1/2 to base into 3–5 oblong to ovate teeth, sometimes entire, sparsely short-strigose to -villous, sometimes glabrate.

Cauline leaves

(0–)1;

blade reduced.

Inflorescences

20–200-flowered, congested, (1–)1.5–6(–10) cm diam.

(5–)8–60-flowered, 2–12 cm diam.

Pedicels

1.5–6 mm.

5–30(–35) mm.

Flowers

4–6 mm diam.;

epicalyx bractlets 5, lanceolate to elliptic, 0.6–1.5(–2) mm;

hypanthium shallowly cupulate, 1 × 2–3 mm;

sepals (1–)1.5–2.5(–3) mm, acute;

petals white to pale yellowish, linear, 1 mm;

stamens 5, filaments 0.3–1 mm, anthers yellow with maroon margins, ovate, 0.3–0.5 mm;

carpels 1–2(–3), styles 0.7–1.8 mm.

9–12 mm diam.;

epicalyx bractlets narrowly lanceolate to lanceolate, 1–2 mm;

hypanthium interior brick- to blood-red or golden, 0.5–1.5 × 3–5 mm;

sepals 3–4.5 mm, base brick- to blood-red adaxially, apex acute to acuminate;

petals brick- to blood-red, linear to oblanceolate or narrowly elliptic, 2–3.5 mm;

stamens 5, filaments 1.3–2 mm, anthers maroon, oblong, 0.6–0.9 mm;

carpels 1–5, styles 2–3 mm.

Achenes

brown, 0.8–1.5 mm, smooth, prominently carunculate.

brown, 2–2.5 mm.

2n

= 28.

Ivesia paniculata

Ivesia multifoliolata

Phenology Flowering summer. Flowering summer.
Habitat Dry shallow volcanic ash and cinders atop volcanic bedrock, open sagebrush communities, adjacent conifer woodlands Dry flats and slopes, on gravelly volcanic or sandstone soil, often along seasonal stream courses, in oak and conifer woodlands
Elevation 1500–1800 m (4900–5900 ft) 1800–2400 m (5900–7900 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
AZ
[BONAP county map]
Discussion

Of conservation concern.

Ivesia paniculata is known only from the Ash Valley area of Lassen County. The distinctions between I. paniculata and I. rhypara are perhaps on the same scale as variation among disjunct population clusters of I. rhypara, but no taxonomic adjustments are proposed at this time.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Ivesia multifoliolata is found between the Grand Canyon and Mogollon Rim in northern Arizona. It is both the only species of the genus with red petals and the only one that overlaps the range of the red-petaled Potentilla sect. Rubrae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 229. FNA vol. 9, p. 244.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Setosae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Comarella
Sibling taxa
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
Synonyms Horkelia multifoliolata, Comarella multifoliolata, Potentilla multifoliolata
Name authority T. W. Nelson & J. P. Nelson: Brittonia 33: 165, fig. 1. (1981) (Torrey) D. D. Keck: Lloydia 1: 125. (1939)
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