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Coconino ivesia, manyleaf mousetail, red comarella

dwarf ivesia, dwarf mousetail

Habit Plants green. Plants green, tufted to ± tightly matted; taproot stout, not fleshy.
Stems

ascending to erect, 2–6 dm.

decumbent to erect, 0.3–1(–1.5) dm.

Basal leaves

6–25(–30) cm; sheathing base minutely glandular abaxially;

petiole 0.5–8(–10) cm;

leaflets 12–30 per side, obovate to broadly cuneate, 5–12(–15) mm, incised 1/3–1/2 to base into 3–5 oblong to ovate teeth, sometimes entire, sparsely short-strigose to -villous, sometimes glabrate.

tightly cylindric, (0.5–)1–8(–12) cm; sheathing base usually ± strigose abaxially;

petiole 0.5–4 cm, hairs 0.2–1 mm;

leaflets 10–15(–20) per side, 0.5–4(–5) mm, sparsely to moderately short-villous, densely glandular, lobes 3–6(–8), oblanceolate to narrowly oblong or oval, apex setose.

Cauline leaves

(0–)1, not paired.

Inflorescences

(5–)8–60-flowered, 2–12 cm diam.

(1–)5–10-flowered, 0.8–3 cm diam.;

glomerules usually 1.

Pedicels

5–30(–35) mm.

1.5–8(–10) mm.

Flowers

9–12 mm diam.;

epicalyx bractlets narrowly lanceolate to lanceolate, 1–2 mm;

hypanthium interior brick- to blood-red or golden, 0.5–1.5 × 3–5 mm;

sepals 3–4.5 mm, base brick- to blood-red adaxially, apex acute to acuminate;

petals brick- to blood-red, linear to oblanceolate or narrowly elliptic, 2–3.5 mm;

stamens 5, filaments 1.3–2 mm, anthers maroon, oblong, 0.6–0.9 mm;

carpels 1–5, styles 2–3 mm.

9–11 mm diam.;

epicalyx bractlets elliptic to oblong, 1.2–2 mm;

hypanthium shallowly cupulate, 1–2 × 2.5–5(–7) mm;

sepals 2–3.5 mm, bluntly acute;

petals golden yellow, broadly oblanceolate to spatulate or broadly obovate, (2–)2.5–3.5(–4) mm;

stamens 10, filaments 0.6–1.8 mm, anthers yellow, 0.6–0.8 mm;

carpels 10–30, styles 0.9–1.3 mm.

Achenes

brown, 2–2.5 mm.

greenish tan to light brown, 1.2–1.5 mm.

2n

= 28.

Ivesia multifoliolata

Ivesia pygmaea

Phenology Flowering summer. Flowering summer.
Habitat Dry flats and slopes, on gravelly volcanic or sandstone soil, often along seasonal stream courses, in oak and conifer woodlands Dry rocky slopes, sometimes in horizontal rock crevices, in high-elevation sagebrush communities, subalpine to alpine conifer woodlands, alpine tundra
Elevation 1800–2400 m (5900–7900 ft) 2700–4000 m (8900–13100 ft)
Distribution
from FNA
AZ
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Ivesia multifoliolata is found between the Grand Canyon and Mogollon Rim in northern Arizona. It is both the only species of the genus with red petals and the only one that overlaps the range of the red-petaled Potentilla sect. Rubrae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Ivesia pygmaea is known only from the southern Sierra Nevada in California, where it overlaps the range of I. lycopodioides var. megalopetala. Stamen number is the only unequivocal way to distinguish between the two, though I. pygmaea is also more generally setose with somewhat shallower hypanthia. It is also more likely to have much-branched caudices and can form mats in rocky sites. Larger plants with relatively open cymes have sometime been segregated as I. chaetophora; this probably represents ecological and phenotypic plasticity.

Early reports of Ivesia pygmaea from Nevada are based on P. A. Rydberg’s (1898) mistaken understanding of the type locality, which is actually in Tulare County, California. A reported occurrence from the northern Sierra Nevada in Nevada (J. T. Kartesz 1987) is presumably based on the same source as the unconfirmed report of I. lycopodioides from the same locality (D. D. Keck 1938).

The correct name for this species if treated as Potentilla is P. nubigena Greene; P. decipiens Greene is a later homonym and illegitimate.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 244. FNA vol. 9, p. 231.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Comarella Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Ivesia
Sibling taxa
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
Synonyms Horkelia multifoliolata, Comarella multifoliolata, Potentilla multifoliolata I. chaetophora, I. gordonii var. pygmaea, Potentilla gordonii var. chaetophora, P. nubigena
Name authority (Torrey) D. D. Keck: Lloydia 1: 125. (1939) A. Gray: Proc. Amer. Acad. Arts 6: 531. (1865)
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