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Coconino ivesia, manyleaf mousetail, red comarella

Pickering's ivesia, silky mousetail

Habit Plants green. Plants grayish green; glands abundant.
Stems

ascending to erect, 2–6 dm.

ascending to erect, 3–5 dm.

Basal leaves

6–25(–30) cm; sheathing base minutely glandular abaxially;

petiole 0.5–8(–10) cm;

leaflets 12–30 per side, obovate to broadly cuneate, 5–12(–15) mm, incised 1/3–1/2 to base into 3–5 oblong to ovate teeth, sometimes entire, sparsely short-strigose to -villous, sometimes glabrate.

8–20 cm; sheathing base ± strigose abaxially;

stipules linear to narrowly lanceolate, 3–5 mm;

petiole 1–3.5 cm, hairs abundant, ascending to spreading, 1–2 mm;

leaflets 35–50 per side, loosely overlapping, 2–6 mm, lobes 3–5, oblanceolate to obovate or oval, hairs abundant, ± ascending, 1–2(–3) mm.

Cauline leaves

5–10.

Inflorescences

(5–)8–60-flowered, 2–12 cm diam.

10–100-flowered, (1.5–)5–15 cm diam., flowers usually arranged individually, sometimes more congested.

Pedicels

5–30(–35) mm.

(1–)2–10 mm.

Flowers

9–12 mm diam.;

epicalyx bractlets narrowly lanceolate to lanceolate, 1–2 mm;

hypanthium interior brick- to blood-red or golden, 0.5–1.5 × 3–5 mm;

sepals 3–4.5 mm, base brick- to blood-red adaxially, apex acute to acuminate;

petals brick- to blood-red, linear to oblanceolate or narrowly elliptic, 2–3.5 mm;

stamens 5, filaments 1.3–2 mm, anthers maroon, oblong, 0.6–0.9 mm;

carpels 1–5, styles 2–3 mm.

8–13 mm diam.;

epicalyx bractlets linear-lanceolate to lanceolate, (1.8–)2–2.5 mm;

hypanthium cupulate to turbinate, 1.5–3 × 2.5–4 mm, often nearly as deep as wide;

sepals often purple-suffused or -mottled, (2.5–)3–5 mm, acuminate;

petals white, becoming pink-tinged with age, oblanceolate to narrowly spatulate, (2.8–)3–5(–6) mm;

stamens 20, filaments filiform, 1.5–2.3 mm, anthers white to cream, 0.3–0.6 mm;

carpels 2–4, styles 2.5–3.5 mm.

Achenes

brown, 2–2.5 mm.

dark brown, 2.5–3 mm.

2n

= 28.

Ivesia multifoliolata

Ivesia pickeringii

Phenology Flowering summer. Flowering summer.
Habitat Dry flats and slopes, on gravelly volcanic or sandstone soil, often along seasonal stream courses, in oak and conifer woodlands Moist, rocky, grassy meadows, mainly on ultramafic-derived clayey soil, in montane conifer woodlands
Elevation 1800–2400 m (5900–7900 ft) 800–1500 m (2600–4900 ft)
Distribution
from FNA
AZ
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Of conservation concern.

Ivesia multifoliolata is found between the Grand Canyon and Mogollon Rim in northern Arizona. It is both the only species of the genus with red petals and the only one that overlaps the range of the red-petaled Potentilla sect. Rubrae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Ivesia pickeringii is known from Siskiyou and Trinity counties. Some early treatments (for example, P. A. Rydberg 1898, 1908–1918; W. L. Jepson [1923–1925], 1909–1943, vol. 2) conflated this species with I. aperta, hence reports of I. pickeringii in the Sierra Nevada. Historic collections from Edgewood, at the headwaters of the Shasta River in Siskiyou County, have inflorescences more glomerulate than elsewhere in the species range.

The chromosome count of 2n = 28 (P. A. Munz 1959) needs confirmation. If based on Kruckeberg 3665, which was distributed as a voucher for that count, the determination of this collection has been changed to Ivesia sericoleuca.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 244. FNA vol. 9, p. 239.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Comarella Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Unguiculatae
Sibling taxa
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
Synonyms Horkelia multifoliolata, Comarella multifoliolata, Potentilla multifoliolata Potentilla pickeringii
Name authority (Torrey) D. D. Keck: Lloydia 1: 125. (1939) Torrey ex A. Gray: Proc. Amer. Acad. Arts 6: 531. (1865)
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