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Coconino ivesia, manyleaf mousetail, red comarella

Charleston Peak ivesia, Charleston Peak mousetail

Habit Plants green. Plants green, diffusely matted.
Stems

ascending to erect, 2–6 dm.

prostrate to decumbent, 0.2–0.6(–1) dm.

Basal leaves

6–25(–30) cm; sheathing base minutely glandular abaxially;

petiole 0.5–8(–10) cm;

leaflets 12–30 per side, obovate to broadly cuneate, 5–12(–15) mm, incised 1/3–1/2 to base into 3–5 oblong to ovate teeth, sometimes entire, sparsely short-strigose to -villous, sometimes glabrate.

tightly cylindric, 1–4(–5) cm; sheathing base mostly not strigose abaxially;

petiole 0.3–2 cm, hairs 0.5–1 mm;

lateral leaflets 5–10(–12) per side, ± overlapping, ± flabellate, 1–2.5 mm, incised to base or nearly so into 3–5 obovate to oval lobes, apex sometimes ± setose, surfaces loosely long-strigose, sparsely glandular;

terminal leaflets indistinct.

Cauline leaves

1(–2);

blade vestigial.

Inflorescences

(5–)8–60-flowered, 2–12 cm diam.

1–10-flowered, ± congested, 0.5–1.5 cm diam.

Pedicels

5–30(–35) mm.

3–7(–10) mm.

Flowers

9–12 mm diam.;

epicalyx bractlets narrowly lanceolate to lanceolate, 1–2 mm;

hypanthium interior brick- to blood-red or golden, 0.5–1.5 × 3–5 mm;

sepals 3–4.5 mm, base brick- to blood-red adaxially, apex acute to acuminate;

petals brick- to blood-red, linear to oblanceolate or narrowly elliptic, 2–3.5 mm;

stamens 5, filaments 1.3–2 mm, anthers maroon, oblong, 0.6–0.9 mm;

carpels 1–5, styles 2–3 mm.

6–9 mm diam.;

epicalyx bractlets 5, lanceolate to narrowly oblong or narrowly ovate, 0.8–1.3 mm;

hypanthium shallowly cupulate, ± 1 × 2–3(–4) mm;

sepals 1.5–2.3 mm, ± acute;

petals yellow, oblanceolate to spatulate, 2–3.2 mm;

stamens 5, filaments 0.9–1.2 mm, anthers yellow, oblong, 0.7–0.9 mm;

carpels 6–10, styles 1–1.5 mm.

Achenes

brown, 2–2.5 mm.

greenish cream to light brown, 1.4–1.8(–2) mm, smooth, not carunculate.

2n

= 28.

Ivesia multifoliolata

Ivesia cryptocaulis

Phenology Flowering summer. Flowering summer.
Habitat Dry flats and slopes, on gravelly volcanic or sandstone soil, often along seasonal stream courses, in oak and conifer woodlands Dry to moist limestone gravel, scree, or talus slopes and outcrops, subalpine conifer woodlands, alpine tundra communities
Elevation 1800–2400 m (5900–7900 ft) 2500–3700 m (8200–12100 ft)
Distribution
from FNA
AZ
[BONAP county map]
from FNA
NV
[BONAP county map]
Discussion

Of conservation concern.

Ivesia multifoliolata is found between the Grand Canyon and Mogollon Rim in northern Arizona. It is both the only species of the genus with red petals and the only one that overlaps the range of the red-petaled Potentilla sect. Rubrae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Ivesia cryptocaulis is known only from timberline and above in the Spring Mountains of Clark County. Plants differ from all others of the genus by having slender, elongate, diffuse caudex branches that ramify through its habitat of gravel, scree, and talus. The species is here provisionally included in sect. Setosae as a diffusely matted relative of I. shockleyi (as also suggested by D. D. Keck 1938); the ecarunculate seeds and relatively straight pedicels may indicate a greater similarity to I. pygmaea in sect. Ivesia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 244. FNA vol. 9, p. 228.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Comarella Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Setosae
Sibling taxa
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
Synonyms Horkelia multifoliolata, Comarella multifoliolata, Potentilla multifoliolata Potentilla cryptocaulis
Name authority (Torrey) D. D. Keck: Lloydia 1: 125. (1939) (Clokey) D. D. Keck: Lloydia 1: 130. (1939)
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