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Coconino ivesia, manyleaf mousetail, red comarella

Bailey's ivesia, Owyhee ivesia

Habit Plants green. Plants green, ± tufted, often forming hanging clumps, sometimes rosetted.
Stems

ascending to erect, 2–6 dm.

pendent or prostrate to ascending, 0.5–2(–2.5) dm.

Basal leaves

6–25(–30) cm; sheathing base minutely glandular abaxially;

petiole 0.5–8(–10) cm;

leaflets 12–30 per side, obovate to broadly cuneate, 5–12(–15) mm, incised 1/3–1/2 to base into 3–5 oblong to ovate teeth, sometimes entire, sparsely short-strigose to -villous, sometimes glabrate.

planar, 3–12(–18) cm; sheathing base not or sparsely strigose abaxially;

petiole 1–8 cm;

lateral leaflets 2–6(–10) per side, separate to slightly overlapping distally, ovate or obovate to flabellate, 4–15(–25) mm, incised 1/4–3/4 to base into 3–11(–15) ovate to oblanceolate teeth or lobes, apex not setose, surfaces ± sparsely short-pilose or hirsute, ± glandular;

terminal leaflets ± distinct.

Cauline leaves

1–2;

blade reduced.

Inflorescences

(5–)8–60-flowered, 2–12 cm diam.

(1–)5–40-flowered, open, (1–)1.5–8(–10) cm diam.

Pedicels

5–30(–35) mm.

2–15(–30) mm.

Flowers

9–12 mm diam.;

epicalyx bractlets narrowly lanceolate to lanceolate, 1–2 mm;

hypanthium interior brick- to blood-red or golden, 0.5–1.5 × 3–5 mm;

sepals 3–4.5 mm, base brick- to blood-red adaxially, apex acute to acuminate;

petals brick- to blood-red, linear to oblanceolate or narrowly elliptic, 2–3.5 mm;

stamens 5, filaments 1.3–2 mm, anthers maroon, oblong, 0.6–0.9 mm;

carpels 1–5, styles 2–3 mm.

4–10 mm diam.;

epicalyx bractlets 5, narrowly lanceolate to ovate, 0.8–2.5 mm;

hypanthium patelliform, 0.5–2 × 2–4 mm;

sepals (1.2–)1.5–4 mm, acute;

petals white or pale yellow, oblanceolate to narrowly spatulate, 1.5–2.5 mm;

stamens 5, filaments 0.5–1.1 mm, anthers yellow, sometimes with reddish margins, oblong, 0.4–0.7 mm;

carpels (1–)3–8, styles 0.9–1.8 mm.

Achenes

brown, 2–2.5 mm.

greenish white to light tan, 1.5–2 mm, smooth or rugose, ± carunculate.

2n

= 28.

Ivesia multifoliolata

Ivesia baileyi

Phenology Flowering summer.
Habitat Dry flats and slopes, on gravelly volcanic or sandstone soil, often along seasonal stream courses, in oak and conifer woodlands
Elevation 1800–2400 m (5900–7900 ft)
Distribution
from FNA
AZ
[BONAP county map]
from FNA
CA; ID; NV; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Ivesia multifoliolata is found between the Grand Canyon and Mogollon Rim in northern Arizona. It is both the only species of the genus with red petals and the only one that overlaps the range of the red-petaled Potentilla sect. Rubrae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

The two varieties accepted here were not recognized by D. D. Keck (1938), who instead treated Ivesia setosa as a variety of I. baileyi. Field investigations confirm that the three entities are reasonably distinct morphologically, with intergradation where their otherwise distinct ranges overlap in central Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals pale yellow; hypanthium interior yellow to golden; epicalyx bractlets ± 3/4 to as long as sepals; achenes smooth.
var. baileyi
1. Petals white; hypanthium interior pale green or cream to maroon; epicalyx bractlets usually less than 1/2 as long as sepals; achenes rugose.
var. beneolens
Source FNA vol. 9, p. 244. FNA vol. 9, p. 226.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Comarella Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Setosae
Sibling taxa
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
I. aperta, I. argyrocoma, I. arizonica, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
Subordinate taxa
I. baileyi var. baileyi, I. baileyi var. beneolens
Synonyms Horkelia multifoliolata, Comarella multifoliolata, Potentilla multifoliolata Potentilla baileyi
Name authority (Torrey) D. D. Keck: Lloydia 1: 125. (1939) S. Watson: Botany (Fortieth Parallel), 90. (1871)
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