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Coconino ivesia, manyleaf mousetail, red comarella

Sierra Valley ivesia, Sierra Valley mousetail

Habit Plants green. Plants greenish gray to silvery; glands sparse.
Stems

ascending to erect, 2–6 dm.

decumbent to erect, 1.5–4.5 dm.

Basal leaves

6–25(–30) cm; sheathing base minutely glandular abaxially;

petiole 0.5–8(–10) cm;

leaflets 12–30 per side, obovate to broadly cuneate, 5–12(–15) mm, incised 1/3–1/2 to base into 3–5 oblong to ovate teeth, sometimes entire, sparsely short-strigose to -villous, sometimes glabrate.

(7–)10–20(–23) cm; sheathing base densely strigose abaxially;

stipules absent;

petiole 4–8 cm, hairs abundant, ascending, ± 1(–3) mm;

leaflets 20–35 per side, loosely overlapping, 3–15(–20) mm, lobes 0–4(–5), oblanceolate to elliptic, hairs abundant, ascending to appressed, ± 1(–2) mm.

Cauline leaves

3–8.

Inflorescences

(5–)8–60-flowered, 2–12 cm diam.

(10–)20–250-flowered, 1–14 cm diam., flowers arranged in several to many tight glomerules of 5–20 flowers.

Pedicels

5–30(–35) mm.

1–3(–15) mm.

Flowers

9–12 mm diam.;

epicalyx bractlets narrowly lanceolate to lanceolate, 1–2 mm;

hypanthium interior brick- to blood-red or golden, 0.5–1.5 × 3–5 mm;

sepals 3–4.5 mm, base brick- to blood-red adaxially, apex acute to acuminate;

petals brick- to blood-red, linear to oblanceolate or narrowly elliptic, 2–3.5 mm;

stamens 5, filaments 1.3–2 mm, anthers maroon, oblong, 0.6–0.9 mm;

carpels 1–5, styles 2–3 mm.

5–12 mm diam.;

epicalyx bractlets lanceolate, 1.5–2.5(–3.5) mm;

hypanthium cupulate, 1–2 × 2.5–4(–5) mm, 1/2–2/3 as deep as wide;

sepals green, 2.5–4(–5.5) mm, acute to acuminate;

petals light yellow, oblanceolate to broadly obovate, 2–7 mm;

stamens 20, filaments filiform, 1–4 mm, anthers yellow, 0.4–0.6 mm;

carpels 2–7, styles 2–4 mm.

Achenes

brown, 2–2.5 mm.

brown, 2–3 mm.

2n

= 28.

Ivesia multifoliolata

Ivesia aperta

Phenology Flowering summer.
Habitat Dry flats and slopes, on gravelly volcanic or sandstone soil, often along seasonal stream courses, in oak and conifer woodlands
Elevation 1800–2400 m (5900–7900 ft)
Distribution
from FNA
AZ
[BONAP county map]
from FNA
CA; NV
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Ivesia multifoliolata is found between the Grand Canyon and Mogollon Rim in northern Arizona. It is both the only species of the genus with red petals and the only one that overlaps the range of the red-petaled Potentilla sect. Rubrae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Many historic collections of Ivesia aperta were identified as I. pickeringii. D. D. Keck (1938) considered I. aperta to be no more than a yellow-flowered form of I. sericoleuca, a situation clarified and rectified by J. T. Howell (1962). The ranges of the two species overlap in Sierra Valley, California, with mixed populations occurring near Beckwourth and in the valleys north of Sierra Valley. In general, var. aperta occurs in the southeastern portions of the valley and I. sericoleuca occurs on the western side; the species are not otherwise sympatric. In addition to petal color, which fades in herbarium material, I. aperta differs from I. sericoleuca in having smaller flowers with shallower hypanthia. Hairs at the base of the stems and petioles of I. aperta are relatively short (to 2 mm) and generally ascending; those of I. sericoleuca are longer (to 4 mm) and spreading. Variety canina combines the petal color of var. aperta and floral dimensions of I. sericoleuca.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals 2–3 mm, oblanceolate; filaments 1–1.5(–2) mm; Sierra Valley drainage, California, Carson and Virginia ranges, Nevada.
var. aperta
1. Petals usually 4–7 mm, narrowly to broadly obovate; filaments 2–4 mm; Dog Valley, California, Nevada.
var. canina
Source FNA vol. 9, p. 244. FNA vol. 9, p. 240.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Comarella Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Unguiculatae
Sibling taxa
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
Subordinate taxa
I. aperta var. aperta, I. aperta var. canina
Synonyms Horkelia multifoliolata, Comarella multifoliolata, Potentilla multifoliolata Potentilla aperta
Name authority (Torrey) D. D. Keck: Lloydia 1: 125. (1939) (J. T. Howell) Munz: Suppl. Calif. Fl., 111. (1968)
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