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club-moss ivesia, clubmoss mousetail

Habit Plants green, usually rosetted, sometimes ± tufted; taproot fusiform, fleshy. Herbs, shrubs, or subshrubs.
Stems

decumbent to erect, 0.3–3 dm.

Leaves

alternate, rarely opposite, pinnately compound, sometimes simple or palmately compound;

stipules present, rarely absent.

Basal leaves

tightly to loosely cylindric, 1–15 cm; sheathing base glabrous abaxially;

petiole 0.5–4 cm, hairs 0.2–1 mm;

leaflets 10–35 per side, 1–8 mm, glabrous or short-hirsute, minutely glandular, lobes (2–)4–8(–10), linear to obovate or ± orbiculate, apex sometimes setose.

Cauline leaves

0–2(–3), not paired.

Inflorescences

3–20(–25)-flowered, (0.5–)1–2.5(–3.5) cm diam.;

glomerules usually 1.

Pedicels

(0.5–)1–7(–11) mm.

Flowers

6–12 mm diam.;

epicalyx bractlets oblong to oval, 0.8–2.5(–3) mm;

hypanthium shallowly cupulate, 1–2 × 2.5–5 mm;

sepals (1.8–)2–4(–4.5) mm, obtuse to acute;

petals golden yellow, obovate, 2–5 mm;

stamens 5, filaments 0.8–2 mm, anthers yellow, 0.6–0.8 mm;

carpels (5–)8–15(–18), styles 1–3 mm.

torus usually enlarged, sometimes small or absent;

carpels 1–260(–450), distinct, free, styles distinct, rarely connate (Roseae);

ovules 1(or 2), collateral (Rubeae) or superposed (Fallugia, Filipendula).

Fruits

achenes or aggregated achenes sometimes with fleshy, urn-shaped hypanthium or enlarged torus, sometimes aggregated drupelets;

styles persistent or deciduous, not elongate (elongate but not plumose in Geum).

Achenes

greenish tan to light brown, 1.2–1.5 mm.

x

= 7(8).

Ivesia lycopodioides

Rosaceae subfam. rosoideae

Distribution
from FNA
CA; NV
[WildflowerSearch map]
[BONAP county map]
North America; Mexico; Central America; South America; West Indies; Bermuda; Eurasia; Africa; Atlantic Islands; Indian Ocean Islands; Pacific Islands; Australia
Discussion

Varieties 3 (3 in the flora).

The three varieties of Ivesia lycopodioides are for the most part readily distinguished, though intergradation is known. The high-elevation var. lycopodioides extends farthest north; var. scandularis is the only variety in the White Mountains. Variety megalopetala is found mostly at somewhat lower (subalpine) elevations and generally has a more southern range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Variation in the number of genera in subfam. Rosoideae is due to differences in generic delimitation between D. Potter et al. (2007) and the authors of some Potentilleae genera. Cyanogenic glycosides and sorbitol are absent in the subfamily.

Tribes 6, genera 28–35, species ca. 1600 (6 tribes, 26 genera, 302 species, including 1 hybrid, in the flora)

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaflets short-hirsute, apical setae (0–)0.5–1(–2) mm; White Mountains and c Sierra Nevada.
var. scandularis
1. Leaflets glabrous or sparsely short-hirsute, apical setae 0–0.5 mm; Sierra Nevada and Sweetwater Mountains
→ 2
2. Leaflet lobes ± orbiculate, ± 1 mm; petals 2–3 × 1 mm.
var. lycopodioides
2. Leaflet lobes linear to oblanceolate, 2–8 mm; petals 3–5 × 2–4 mm.
var. megalopetala
Source FNA vol. 9, p. 232. FNA vol. 9, p. 23. Author: Luc Brouillet.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Ivesia Rosaceae
Sibling taxa
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
Subordinate taxa
I. lycopodioides var. lycopodioides, I. lycopodioides var. megalopetala, I. lycopodioides var. scandularis
Synonyms Potentilla lycopodioides
Name authority A. Gray: Proc. Amer. Acad. Arts 6: 530. (1865) Arnott: Botany, 107. (1832)
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