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club-moss ivesia, clubmoss mousetail

grimy ivesia, grimy mousetail

Habit Plants green, usually rosetted, sometimes ± tufted; taproot fusiform, fleshy. Plants grayish, ± matted.
Stems

decumbent to erect, 0.3–3 dm.

± prostrate, (0.1–)0.2–1.5(–2) dm.

Basal leaves

tightly to loosely cylindric, 1–15 cm; sheathing base glabrous abaxially;

petiole 0.5–4 cm, hairs 0.2–1 mm;

leaflets 10–35 per side, 1–8 mm, glabrous or short-hirsute, minutely glandular, lobes (2–)4–8(–10), linear to obovate or ± orbiculate, apex sometimes setose.

tightly cylindric, 2–10 cm; sheathing base densely hairy abaxially;

petiole 0.5–4(–10) cm;

lateral leaflets 4–10(–15) per side, overlapping at least distally, ± flabellate, 0.5–3(–4) mm, incised to base or nearly so into (0–)2–4(–9) elliptic to obovate or orbiculate lobes, apex usually not setose, surfaces densely hirsute, cryptically glandular;

terminal leaflets indistinct.

Cauline leaves

0–2(–3), not paired.

(0–)1;

blade vestigial.

Inflorescences

3–20(–25)-flowered, (0.5–)1–2.5(–3.5) cm diam.;

glomerules usually 1.

5–60(–100)-flowered, congested, 0.5–5(–7) cm diam.

Pedicels

(0.5–)1–7(–11) mm.

1.5–4(–8) mm.

Flowers

6–12 mm diam.;

epicalyx bractlets oblong to oval, 0.8–2.5(–3) mm;

hypanthium shallowly cupulate, 1–2 × 2.5–5 mm;

sepals (1.8–)2–4(–4.5) mm, obtuse to acute;

petals golden yellow, obovate, 2–5 mm;

stamens 5, filaments 0.8–2 mm, anthers yellow, 0.6–0.8 mm;

carpels (5–)8–15(–18), styles 1–3 mm.

4–8 mm diam.;

epicalyx bractlets 5, linear to elliptic or narrowly ovate, 0.8–1.6(–2.1) mm;

hypanthium patelliform to shallowly cupulate, 1–1.5 × (1.5–)2–3(–3.5) mm;

sepals 1.4–2.5(–2.8) mm, acute;

petals white to pale yellowish, linear to narrowly oblanceolate, 1–1.5 mm;

stamens 5, filaments 0.6–1.8 mm, anthers maroon or yellow with maroon margins, oblong, 0.3–0.6 mm;

carpels 1–2(–4), styles 1–1.5 mm.

Achenes

greenish tan to light brown, 1.2–1.5 mm.

brown, 1.2–1.6(–2) mm, smooth, prominently carunculate.

Ivesia lycopodioides

Ivesia rhypara

Distribution
from FNA
CA; NV
[WildflowerSearch map]
[BONAP county map]
from FNA
NV; OR
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[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

The three varieties of Ivesia lycopodioides are for the most part readily distinguished, though intergradation is known. The high-elevation var. lycopodioides extends farthest north; var. scandularis is the only variety in the White Mountains. Variety megalopetala is found mostly at somewhat lower (subalpine) elevations and generally has a more southern range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Ivesia rhypara forms densely hairy mats or mounds on isolated edaphic sites in the northern Great Basin. Only one of the widely disjunct population clusters is recognized as a distinct variety; the others have general differences in gestalt, but no significant distinctions have been determined (B. Ertter 1989). Isozyme analysis likewise does not show significant differences (T. Kaye et al., unpubl., BLM and Oregon Dept. of Agriculture, 1991). Reproductive studies by D. Wiens (pers. comm.) show low seed set (ca. 5% in var. rhypara, somewhat higher in var. shellyi), possibly indicating a deleterious genetic load.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaflets short-hirsute, apical setae (0–)0.5–1(–2) mm; White Mountains and c Sierra Nevada.
var. scandularis
1. Leaflets glabrous or sparsely short-hirsute, apical setae 0–0.5 mm; Sierra Nevada and Sweetwater Mountains
→ 2
2. Leaflet lobes ± orbiculate, ± 1 mm; petals 2–3 × 1 mm.
var. lycopodioides
2. Leaflet lobes linear to oblanceolate, 2–8 mm; petals 3–5 × 2–4 mm.
var. megalopetala
1. Plants (4–)8–30 cm diam.; inflorescences (1–)2–5(–7) cm diam.; scarcely or cryptically petrophytic in bedrock overlain by rubble and soil.
var. rhypara
1. Plants 2–10(–15) cm diam.; inflorescences 0.5–2(–5) cm diam.; overtly petrophytic in boulders and outcrops.
var. shellyi
Source FNA vol. 9, p. 232. FNA vol. 9, p. 228.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Ivesia Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Setosae
Sibling taxa
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
Subordinate taxa
I. lycopodioides var. lycopodioides, I. lycopodioides var. megalopetala, I. lycopodioides var. scandularis
I. rhypara var. rhypara, I. rhypara var. shellyi
Synonyms Potentilla lycopodioides
Name authority A. Gray: Proc. Amer. Acad. Arts 6: 530. (1865) Ertter & Reveal: Madroño 24: 224, fig. 1. (1977)
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