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club-moss ivesia, clubmoss mousetail

Pickering's ivesia, silky mousetail

Habit Plants green, usually rosetted, sometimes ± tufted; taproot fusiform, fleshy. Plants grayish green; glands abundant.
Stems

decumbent to erect, 0.3–3 dm.

ascending to erect, 3–5 dm.

Basal leaves

tightly to loosely cylindric, 1–15 cm; sheathing base glabrous abaxially;

petiole 0.5–4 cm, hairs 0.2–1 mm;

leaflets 10–35 per side, 1–8 mm, glabrous or short-hirsute, minutely glandular, lobes (2–)4–8(–10), linear to obovate or ± orbiculate, apex sometimes setose.

8–20 cm; sheathing base ± strigose abaxially;

stipules linear to narrowly lanceolate, 3–5 mm;

petiole 1–3.5 cm, hairs abundant, ascending to spreading, 1–2 mm;

leaflets 35–50 per side, loosely overlapping, 2–6 mm, lobes 3–5, oblanceolate to obovate or oval, hairs abundant, ± ascending, 1–2(–3) mm.

Cauline leaves

0–2(–3), not paired.

5–10.

Inflorescences

3–20(–25)-flowered, (0.5–)1–2.5(–3.5) cm diam.;

glomerules usually 1.

10–100-flowered, (1.5–)5–15 cm diam., flowers usually arranged individually, sometimes more congested.

Pedicels

(0.5–)1–7(–11) mm.

(1–)2–10 mm.

Flowers

6–12 mm diam.;

epicalyx bractlets oblong to oval, 0.8–2.5(–3) mm;

hypanthium shallowly cupulate, 1–2 × 2.5–5 mm;

sepals (1.8–)2–4(–4.5) mm, obtuse to acute;

petals golden yellow, obovate, 2–5 mm;

stamens 5, filaments 0.8–2 mm, anthers yellow, 0.6–0.8 mm;

carpels (5–)8–15(–18), styles 1–3 mm.

8–13 mm diam.;

epicalyx bractlets linear-lanceolate to lanceolate, (1.8–)2–2.5 mm;

hypanthium cupulate to turbinate, 1.5–3 × 2.5–4 mm, often nearly as deep as wide;

sepals often purple-suffused or -mottled, (2.5–)3–5 mm, acuminate;

petals white, becoming pink-tinged with age, oblanceolate to narrowly spatulate, (2.8–)3–5(–6) mm;

stamens 20, filaments filiform, 1.5–2.3 mm, anthers white to cream, 0.3–0.6 mm;

carpels 2–4, styles 2.5–3.5 mm.

Achenes

greenish tan to light brown, 1.2–1.5 mm.

dark brown, 2.5–3 mm.

Ivesia lycopodioides

Ivesia pickeringii

Phenology Flowering summer.
Habitat Moist, rocky, grassy meadows, mainly on ultramafic-derived clayey soil, in montane conifer woodlands
Elevation 800–1500 m (2600–4900 ft)
Distribution
from FNA
CA; NV
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

The three varieties of Ivesia lycopodioides are for the most part readily distinguished, though intergradation is known. The high-elevation var. lycopodioides extends farthest north; var. scandularis is the only variety in the White Mountains. Variety megalopetala is found mostly at somewhat lower (subalpine) elevations and generally has a more southern range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Ivesia pickeringii is known from Siskiyou and Trinity counties. Some early treatments (for example, P. A. Rydberg 1898, 1908–1918; W. L. Jepson [1923–1925], 1909–1943, vol. 2) conflated this species with I. aperta, hence reports of I. pickeringii in the Sierra Nevada. Historic collections from Edgewood, at the headwaters of the Shasta River in Siskiyou County, have inflorescences more glomerulate than elsewhere in the species range.

The chromosome count of 2n = 28 (P. A. Munz 1959) needs confirmation. If based on Kruckeberg 3665, which was distributed as a voucher for that count, the determination of this collection has been changed to Ivesia sericoleuca.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaflets short-hirsute, apical setae (0–)0.5–1(–2) mm; White Mountains and c Sierra Nevada.
var. scandularis
1. Leaflets glabrous or sparsely short-hirsute, apical setae 0–0.5 mm; Sierra Nevada and Sweetwater Mountains
→ 2
2. Leaflet lobes ± orbiculate, ± 1 mm; petals 2–3 × 1 mm.
var. lycopodioides
2. Leaflet lobes linear to oblanceolate, 2–8 mm; petals 3–5 × 2–4 mm.
var. megalopetala
Source FNA vol. 9, p. 232. FNA vol. 9, p. 239.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Ivesia Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Unguiculatae
Sibling taxa
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
Subordinate taxa
I. lycopodioides var. lycopodioides, I. lycopodioides var. megalopetala, I. lycopodioides var. scandularis
Synonyms Potentilla lycopodioides Potentilla pickeringii
Name authority A. Gray: Proc. Amer. Acad. Arts 6: 530. (1865) Torrey ex A. Gray: Proc. Amer. Acad. Arts 6: 531. (1865)
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