Ivesia longibracteata |
Rosaceae tribe Potentilleae |
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Castle Crags ivesia, longbract mousetail |
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Habit | Plants green, ± tufted; taproot ± stout, not fleshy. | Herbs, perennial, rarely annual or biennial, shrubs, or subshrubs; unarmed. | ||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | ascending to erect, 0.3–1.2 dm. |
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Leaves | alternate, rarely opposite, pinnately (palmately) compound (simple in Alchemilla, Aphanes, and Chamaerhodos); stipules persistent (absent in Chamaerhodos), adnate to petiole; venation pinnate or palmate. |
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Basal leaves | weakly planar to loosely ± cylindric, (0.5–)2–4(–6) cm; sheathing base glandular abaxially, otherwise glabrous; petiole 0.5–2 cm, hairs 0.2–1 mm; leaflets 5–6 per side, 2–6 mm, ± short-hirsute, glandular-puberulent, lobes 2–7, oblanceolate to spatulate or obovate, apex not setose. |
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Cauline leaves | 1–3, not paired. |
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Inflorescences | 3–14-flowered, 1–2.5 cm diam.; glomerules 1. |
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Pedicels | 1.5–6 mm. |
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Flowers | 8–10 mm diam.; epicalyx bractlets linear to narrowly lanceolate or elliptic, 2.5–5 mm, longer than sepals; hypanthium shallowly cupulate, 0.5–1 × 2–3 mm; sepals 1.5–2.5 mm, ± acute; petals pale yellow, linear to narrowly oblanceolate, 1.5–2.5 mm; stamens 5, filaments 0.7–1.3 mm, anthers yellow, 0.5–0.8 mm; carpels 6–11, styles 1–1.5 mm. |
perianth and androecium perigynous; epicalyx bractlets present, sometimes absent; hypanthium usually patelliform, cupulate, or campanulate, sometimes turbinate, saucer-shaped, flat-bottomed, or subglobose to ellipsoid or ovoid; torus flat to conic or turbinate, enlarged (absent or reduced in Alchemilla, Aphanes, and Chamaerhodos); carpels 1–260, styles basal or lateral to subterminal, distinct; ovules 1(or 2), basal. |
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Fruits | aggregated achenes (achenes in Alchemilla and Aphanes); torus sometimes fleshy; styles deciduous or persistent, not elongate. |
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Achenes | cream to light tan, 1.2–1.5 mm. |
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Ivesia longibracteata |
Rosaceae tribe Potentilleae |
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Phenology | Flowering summer. | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Dry crevices of granodioritic igneous rock, in mixed oak-conifer woodlands | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 1200–1400 m (3900–4600 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
CA |
North America; Mexico; Central America; South America; West Indies; Bermuda; Eurasia; Africa; Atlantic Islands; Pacific Islands; Australia |
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Discussion | Of conservation concern. Ivesia longibracteata is known only from the Castle Crags area of Shasta County. The epithet alludes to a diagnostic characteristic unique in the genus: the epicalyx bractlets are longer than the sepals. The plants grow on vertical rock faces, a habitat more characteristic of sect. Setosae; however, the stems are ascending to erect and do not form hanging clumps, and the pedicels are not curved in fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 14–22, species ca. 860 (14 genera, 189 species, including 1 hybrid, in the flora area). The base chromosome number for Potentilleae is mostly x = 7 (8 in Alchemilla and Aphanes; 14 in Comarum). Variation in the number of genera recognized in Potentilleae is due to differences in generic delimitation between D. Potter et al. (2007) and the authors of Potentilla and segregates here (see 9. Ivesia and 8. Potentilla for discussion). In the former, Duchesnea, Horkelia, Horkeliella, and Ivesia are included within Potentilla. Likewise, Aphanes is included within Alchemilla by Potter et al. while it is kept distinct here. Potentilla and its segregates and Fragaria are host to Phragmidium rusts, but not the other genera of the tribe. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 236. | FNA vol. 9, p. 119. | ||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Ivesia | Rosaceae > subfam. Rosoideae | ||||||||||||||||||||||||||||||||||||||||||||||||||||
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Name authority | Ertter: Syst. Bot. 14: 233, fig. 1. (1989) | Sweet: Brit. Fl. Gard. 2: sub plate 124. (1825) | ||||||||||||||||||||||||||||||||||||||||||||||||||||
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