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Ivesia gordonii

alpine ivesia, alpine mousetail, Gordon's ivesia

Ash Meadows ivesia, King's ivesia, King's mousetail

Habit Plants green, ± tufted, sometimes rosetted; taproot stout, not fleshy. Plants usually grayish green to silvery, often glaucous; glands sparse.
Stems

prostrate to erect, (0.2–)0.5–4 dm.

usually prostrate-decumbent to ascending, sometimes ± erect, 1–4(–5.5) dm.

Basal leaves

tightly to loosely cylindric, (1–)3–20(–25) cm; sheathing base ± glandular abaxially, otherwise glabrous;

petiole 0.5–8 cm, hairs 0.2–0.5 mm;

leaflets (6–)10–25 per side, (1–)2–13(–18) mm, glabrous or ± hirsute or villous, glandular-puberulent or -pubescent, lobes (2–)4–8(–15), linear or narrowly oblanceolate to obovate, apex rarely setose.

5–20 cm; sheathing base glabrous or densely strigose abaxially;

stipules absent or linear to lanceolate, 1–4 mm;

petiole 0.2–1.2 cm, hairs absent or sparse to dense, appressed or ascending, 0.5–2 mm;

leaflets 15–60 per side, loosely to tightly overlapping, (1.8–)2–6(–8) mm, lobes (0–)2–4, narrowly oblanceolate to obovate, hairs absent or sparse to dense, ± appressed, 0.2–0.5(–1) mm.

Cauline leaves

1(–2), not paired.

4–15.

Inflorescences

5–50(–70)-flowered, 1–8(–11) cm diam.;

glomerules 1–several.

5–100-flowered, 1–12 cm diam., flowers arranged individually and/or in few to several loose glomerules of 2–10 flowers.

Pedicels

1–3(–5) mm.

(1–)2–20(–25) mm.

Flowers

5–12 mm diam.;

epicalyx bractlets linear to narrowly elliptic, (0.5–)1–3.5(–4) mm;

hypanthium turbinate to campanulate, (1.5–)2–4(–4.5) × 2–4(–5) mm;

sepals (2–)2.5–5(–6) mm, obtuse to ± acute;

petals yellow, narrowly oblanceolate to narrowly spatulate, (1–)1.5–3 mm;

stamens 5, filaments 1.3–2.5 mm, anthers yellow, sometimes red-margined, 0.5–1 mm;

carpels (1–)2–4(–6), styles 2.5–4.5(–6) mm.

8–12 mm diam.;

epicalyx bractlets lanceolate to elliptic or ovate, (0.8–)1–2(–2.5) mm;

hypanthium shallowly cupulate, 0.5–2 × 1.5–4 mm, ± 1/2 as deep as wide;

sepals sometimes purple-suffused, 2–4(–5) mm, narrowly acute to acuminate;

petals white, spatulate or obovate to orbiculate, (2.8–)3–5(–6) mm;

stamens 20, filaments filiform, 1.5–2.5 mm, anthers white to cream, 0.3–0.6 mm;

carpels 2–9, styles 2–2.5(–3) mm.

Achenes

grayish brown to mottled brown, ± 2 mm.

light brown, 1.8–2.5 mm.

Ivesia gordonii

Ivesia kingii

Distribution
from FNA
CA; CO; ID; MT; NV; OR; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

Ivesia gordonii is the most widespread species of the genus, occurring from Washington to Montana, south to central California and Colorado. The species can be distinguished from other members of sect. Ivesia by the relatively deep hypanthium, which is about as deep as wide. Four varieties are provisionally recognized here (B. Ertter and J. L. Reveal 2007), with the likelihood that future work may indicate additional and/or alternate circumscriptions. Populations that are difficult to assign to a variety can be found where the recognizable taxonomic units come together, for example, in northeastern Utah and western Wyoming, involving var. gordonii and var. wasatchensis, and in California and central Idaho where var. alpicola and var. ursinorum tend to merge.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Significant variation in habit, indument, leaflet lobing, and compactness of inflorescence can be found among and within populations of Ivesia kingii; no taxonomic structure has yet been discerned beyond the varieties recognized here. The variation within widespread var. kingii suggests that var. eremica resulted from a southward, late Pleistocene migration of a hairy phase of I. kingii with compact leaflets out of the Great Basin onto the northern edge of the Mojave Desert (J. L. Reveal 1980). Although densely hairy plants are encountered widely in the Great Basin, only in the Ash Meadows area of southernmost Nye County, the home of var. eremica, is there a consistently hairy phase. The leaflets of var. eremica are so tightly compacted that individual leaflets cannot be distinguished; in var. kingii, leaflets are individually distinct, though plants from Mineral County sometimes approach var. eremica in this regard. In the field, shoots of var. kingii arise directly from a taproot with the spreading branches flowing outwardly from a single point of attachment. In var. eremica, older, mature plants form compact mats with short, spreading caudex branches.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaflets 7–13(–18) mm, glabrous or sparsely hirsute or villous marginally; stems (1–)1.5–4 dm; inflorescences usually branched, 2–8(–11) cm diam., glomerules (1–)2–6(–10), ± capitate.
var. wasatchensis
1. Leaflets (1–)2–7(–9) mm, hairy or ± glabrous except marginally ciliate; stems (0.2–)0.5–2.5 dm; inflorescences simple or branched, 1–3(–5) cm diam., glomerules 1(–3), ± capitate to loosely congested
→ 2
2. Stems usually prostrate to ascending, rarely nearly erect, usually dark reddish, minutely glandular or glandular-puberulent to -pubescent; leaves to 0.8 cm diam.; anthers usually red-margined.
var. ursinorum
2. Stems usually ascending to erect, sometimes decumbent, usually greenish, rarely reddish, hirsute to villous, glandular-puberulent to -pubescent; leaves to 1.5 cm diam.; anthers rarely red-margined
→ 3
3. Stems usually hirsute to villous, sometimes densely so, glandular-pubescent or eglandular; basal leaves (3–)5–10(–15) cm; flowers 7–12 mm diam.; e Idaho and w Montana to Utah, Wyoming, and w Colorado.
var. gordonii
3. Stems not or sparsely hirsute to villous, glandular-puberulent or -pubescent; basal leaves 2–8(–10) cm; flowers 5–9 mm diam.; s Washington to California and e to w Montana.
var. alpicola
1. Leaflets individually ± distinguishable, loosely to tightly overlapping; caudices simple or few branched; Great Basin, ec California, Nevada, w Utah.
var. kingii
1. Leaflets mostly not individually distinguishable, tightly overlapping; caudices often much branched; n Mojave Desert, Nevada.
var. eremica
Source FNA vol. 9, p. 233. FNA vol. 9, p. 238.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Ivesia Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Unguiculatae
Sibling taxa
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
Subordinate taxa
I. gordonii var. alpicola, I. gordonii var. gordonii, I. gordonii var. ursinorum, I. gordonii var. wasatchensis
I. kingii var. eremica, I. kingii var. kingii
Synonyms Horkelia gordonii, Potentilla gordonii Potentilla kingii
Name authority (Hooker) Torrey & A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 6(3): 72. (1858) S. Watson: Botany (Fortieth Parallel), 91, 448. (1871)
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