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purpusia, rock whitefeather

Coconino ivesia, manyleaf mousetail, red comarella

Habit Plants green, ± tufted, often forming hanging clumps, sometimes rosetted. Plants green.
Stems

pendent or prostrate to ascending, (0.2–)0.5–1.6(–3) dm.

ascending to erect, 2–6 dm.

Basal leaves

planar, 2–15(–20) cm; sheathing base usually not strigose abaxially;

petiole 3–10 cm;

lateral leaflets (2–)3–4(–5) per side, separate, broadly ovate or obovate to orbiculate, (2–)5–15(–18) mm, incised 1/4–3/4 to base into (3–)7–11 ± ovate teeth, apex not setose, surfaces ± sparsely short-pilose, ± glandular;

terminal leaflets ± distinct.

6–25(–30) cm; sheathing base minutely glandular abaxially;

petiole 0.5–8(–10) cm;

leaflets 12–30 per side, obovate to broadly cuneate, 5–12(–15) mm, incised 1/3–1/2 to base into 3–5 oblong to ovate teeth, sometimes entire, sparsely short-strigose to -villous, sometimes glabrate.

Cauline leaves

1–3;

blade well developed.

Inflorescences

(1–)5–30(–150)-flowered, open, 0.5–14 cm diam.

(5–)8–60-flowered, 2–12 cm diam.

Pedicels

5–30 mm.

5–30(–35) mm.

Flowers

6–12 mm diam.;

epicalyx bractlets 0(–3), lanceolate, 0.9–1.8 mm;

hypanthium turbinate or campanulate, 1.5–3(–5) × 1–3(–4) mm;

sepals 2–4(–5) mm, acute;

petals yellow or white, oblanceolate to elliptic or obovate, (1.6–)2–4 mm;

stamens 5, filaments 0.8–1.3 mm, anthers ± yellow, oblong to narrowly ovate, 0.6–1.5 mm;

carpels (2–)6–10(–13), atop a stipelike torus, styles 0.9–1.8 mm.

9–12 mm diam.;

epicalyx bractlets narrowly lanceolate to lanceolate, 1–2 mm;

hypanthium interior brick- to blood-red or golden, 0.5–1.5 × 3–5 mm;

sepals 3–4.5 mm, base brick- to blood-red adaxially, apex acute to acuminate;

petals brick- to blood-red, linear to oblanceolate or narrowly elliptic, 2–3.5 mm;

stamens 5, filaments 1.3–2 mm, anthers maroon, oblong, 0.6–0.9 mm;

carpels 1–5, styles 2–3 mm.

Achenes

greenish white to light tan, 1.2–2 mm, faintly rugose, ± carunculate.

brown, 2–2.5 mm.

2n

= 28.

Ivesia arizonica

Ivesia multifoliolata

Phenology Flowering summer.
Habitat Dry flats and slopes, on gravelly volcanic or sandstone soil, often along seasonal stream courses, in oak and conifer woodlands
Elevation 1800–2400 m (5900–7900 ft)
Distribution
from FNA
AZ; CA; NV; UT
[BONAP county map]
from FNA
AZ
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Because the epithet saxosa was retained for Potentilla saxosa upon transfer to Ivesia, a different epithet (arizonica) was needed when Purpusia was likewise transferred in the same publication (B. Ertter 1989). The correct name for this species in Potentilla is P. osterhoutii (A. Nelson) J. T. Howell, due to the existence of P. arizonica.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Ivesia multifoliolata is found between the Grand Canyon and Mogollon Rim in northern Arizona. It is both the only species of the genus with red petals and the only one that overlaps the range of the red-petaled Potentilla sect. Rubrae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals yellow; hypanthia campanulate.
var. arizonica
1. Petals white; hypanthia turbinate.
var. saxosa
Source FNA vol. 9, p. 224. FNA vol. 9, p. 244.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Setosae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Ivesia > sect. Comarella
Sibling taxa
I. aperta, I. argyrocoma, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. multifoliolata, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
I. aperta, I. argyrocoma, I. arizonica, I. baileyi, I. callida, I. campestris, I. cryptocaulis, I. gordonii, I. jaegeri, I. kingii, I. longibracteata, I. lycopodioides, I. muirii, I. paniculata, I. patellifera, I. pickeringii, I. pityocharis, I. pygmaea, I. rhypara, I. sabulosa, I. santolinoides, I. saxosa, I. sericoleuca, I. setosa, I. shockleyi, I. tweedyi, I. unguiculata, I. utahensis, I. webberi
Subordinate taxa
I. arizonica var. arizonica, I. arizonica var. saxosa
Synonyms Purpusia arizonica Horkelia multifoliolata, Comarella multifoliolata, Potentilla multifoliolata
Name authority (Eastwood ex J. T. Howell) Ertter: Syst. Bot. 14: 233. (1989) (Torrey) D. D. Keck: Lloydia 1: 125. (1939)
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