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dune marsh-elder, seacoast marsh elder, seacoast sumpweed

marsh-elder, poverty-weed

Habit Perennials or subshrubs, 30–50(–100) cm. Annuals, perennials, subshrubs, or shrubs, 10–350 cm.
Stems

± decumbent to erect.

usually erect, sometimes decumbent to sprawling, often freely branched.

Leaves

petioles 0–2 mm;

blades narrowly lance-elliptic to lanceolate or spatulate, 1–3(–6+) cm × 4–10(–20) mm, margins usually entire, rarely toothed, faces glabrous, rarely gland-dotted.

cauline; mostly opposite (distal sometimes alternate); petiolate or sessile;

blades (1- or 3-nerved) deltate, elliptic, filiform, lanceolate, linear, obovate, ovate, spatulate, or trullate, margins entire or toothed, faces glabrous or ± scabrellous, often gland-dotted.

Peduncles

2–7+ mm.

Involucres

± campanulate to urceolate, 5–7+ mm.

± hemispheric or turbinate to campanulate or urceolate, 2–10+ mm diam.

Receptacles

flat or convex to hemispheric;

paleae cuneiform or spatulate to linear or setiform, ± membranous, sometimes all or partially wanting.

Pistillate florets

2–5;

corollas 0.3–1.5 mm.

usually 1–8+, rarely 0;

corollas whitish (inconspicuous), ± tubular.

Phyllaries

outer 6–10 distinct, ± herbaceous.

persistent, 3–15+ in 1–3+ series, distinct or ± connate, all ± herbaceous or inner scarious to membranous.

Heads

in racemiform arrays.

discoid or ± disciform, in (± bracteate) racemiform or spiciform arrays (heads 1–2 in axil of each bract, bracts sometimes leaflike).

Cypselae

4–5 mm.

plumply obovoid to pyriform, often ± obcompressed, sometimes scabrellous or hispidulous distally, usually ± gland-dotted;

pappi 0.

Paleae

linear to spatulate (distally dilated), 5–6 mm.

Functionally

staminate florets 8–12+;

corollas 4–5 mm.

staminate florets 3–20+;

corollas whitish to pinkish, funnelform, lobes 5, soon reflexed (filaments ± connate, anthers coherent or distinct).

x

= 18.

2n

= 34.

Iva imbricata

Iva

Phenology Flowering Jul–Oct.
Habitat Dunes, beaches, at or near tidelands
Elevation 0–10 m (0–0 ft)
Distribution
from FNA
AL; FL; GA; LA; MS; NC; SC; TX; VA; West Indies (Bahamas, Cuba)
[WildflowerSearch map]
[BONAP county map]
from USDA
Mostly temperate North America; some subtropical; some introduced in Old World
[BONAP county map]
Discussion

Species ca. 9 (7 in the flora).

M. Bolick (1983), P. O. Karis (1995), and B. Miao et al. (1995, 1995b, 1995c) considered Iva in the sense of R. C. Jackson (1960) to include species that have closer relatives outside Iva in the broad sense than within. I agree and treat the extradited species in Chorisiva, Cyclachaena, Hedosyne, Leuciva, and Oxytenia.

Ivas are wind pollinated, are related to the ragweeds (Ambrosia spp.), and may be similarly associated with pollen allergies.

Records of Iva asperifolia Lessing from Florida are evidently based on specimens that are treated as members of I. angustifolia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Annuals (seldom persisting); leaf blades deltate, filiform, lanceolate, lance-linear, or ovate (rarely succulent or fleshy); heads in ± spiciform arrays; peduncles mostly 0.5–1 mm
→ 2
1. Perennials, subshrubs, or shrubs; leaf blades elliptic, lanceolate, oblanceolate, obovate, ovate, or spatulate (usually ± succulent or fleshy); heads in ± racemiform arrays; peduncles mostly 1–7 mm
→ 4
2. Leaves petiolate, blades mostly deltate to lanceolate, margins toothed
I. annua
2. Leaves ± sessile, blades mostly lance-linear to filiform, margins entire
→ 3
3. Involucres ± prismatically turbinate, 2.5–3 mm; outer phyllaries ± connate
I. angustifolia
3. Involucres ± hemispheric, 1.5–2 mm; outer phyllaries distinct
I. microcephala
4. Subshrubs or shrubs, 50–350 cm; leaf blades ovate or elliptic to lanceolate, margins usually toothed
I. frutescens
4. Perennials or subshrubs, 10–50(–100) cm; leaf blades mostly obovate or elliptic to lanceolate, oblanceolate, or spatulate, margins usually entire, rarely toothed
→ 5
5. Leaf blades glabrous; involucres campanulate to urceolate, 5–7+ mm; cypselae 4–5 mm
I. imbricata
5. Leaf blades usually strigose to scabrellous; involucres ± hemispheric, 2–3.5 mm; cypselae 2–3 mm
→ 6
6. Aerial stems usually woody at bases; outer phyllaries distinct
I. hayesiana
6. Aerial stems usually herbaceous; outer phyllaries ± connate
I. axillaris
Source FNA vol. 21, p. 26. FNA vol. 21, p. 25. Author: John L. Strother.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Ambrosiinae > Iva Asteraceae > tribe Heliantheae > subtribe Ambrosiinae
Sibling taxa
I. angustifolia, I. annua, I. axillaris, I. frutescens, I. hayesiana, I. microcephala
Subordinate taxa
I. angustifolia, I. annua, I. axillaris, I. frutescens, I. hayesiana, I. imbricata, I. microcephala
Name authority Walter: Fl. Carol., 232. (1788) Linnaeus: Sp. Pl. 2: 988. (1753): Gen. Pl. ed. 5, 426. (1754)
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