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isothecium moss, slender mouse-tail moss, tree moss, variable moss

Habit Plants medium-sized, yellowish to dark green, not strongly glossy. Plants small to large, pale to dark green or golden.
Stem(s)

with stipe short to absent, secondary stems 2–4 cm, not julaceous when dry, irregularly branched, sometimes close, branches arching downward, branches of secondary system sharp-tipped, flagelliform branches frequently present, 2–5 cm, attenuate, with narrower smaller leaves;

pseudoparaphyllia foliose, sometimes toothed, often blunt and broad.

and branch leaves often differentiated, loosely appressed, usually ovate, usually concave;

base decurrent;

costa single or double, short, sometimes weak on stipe and stem leaves and strong on branch leaves;

alar cells usually well differentiated, usually quadrate, often excavate;

medial laminal cells short to linear, smooth or prorulose.

Seta

1–2 cm.

long or sometimes short, smooth or roughened throughout or distally.

Sexual condition

dioicous.

Capsule

1–1.5 mm.

erect or sometimes horizontal, cylindric, symmetric or sometimes curved;

annulus usually differentiated;

operculum conic to short-rostrate, straight;

peristome double, perfect or sometimes reduced;

exostome, when not reduced, with external surface cross striolate proximally, papillose distally, when reduced, striate, papillose, or smooth;

endostome basal membrane high or low, segments long, narrowly perforate, cilia nodose or appendiculate.

Calyptra

cucullate, smooth or slightly roughened, naked or hairy.

Primary

stem leaves broadly or narrowly triangular;

margins nearly entire;

apex sometimes attenuate; ecostate or costa present;

alar cells barely differentiated, region small, to 1/10 leaf length.

Branchlet

leaves ovate to ovate-lanceolate;

margins toothed throughout, most strongly in apiculus;

apex attenuate;

costa to 2/3 leaf length;

alar cells usually shorter, except in shoot decurrent portion, region well defined, small, sometimes excavate, at marginal insertion.

Isothecium myosuroides

Lembophyllaceae

Habitat Rock, cliffs, boulders, shaded sites in forests, epiphytic, tree bases
Elevation low to moderate elevations
Distribution
from FNA
AK; ME; NC; NH; TN; NF; NS; PE; Europe; n Africa; Atlantic Islands
[WildflowerSearch map]
North America; Europe; Asia; Atlantic Islands; Pacific Islands
Discussion

The eastern representation of Isothecium myosuroides is clearly the same as the European species, but the relationship of western North American material is unclear. K. Ryall et al. (2005) supported this conclusion with molecular data, but some California specimens identified as this species suggest that further analysis is necessary to clarify their identity. Isothecium myosuroides, although showing some variability, is far more uniform than I. stoloniferum, and the infrequency of sporophytes in the former suggests that cloned populations are frequent. The stems of I. myosuroides are creeping and radiculose.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 14, species ca. 50 (3 genera, 7 species in the flora).

Lembophyllaceae are largely a south-temperate family, but also extend into the tropics. The plants tend to have terete-foliate stems with ovate, concave leaves. The costa is variable, sometimes on different leaves of the same plant. Many of the genera have perfect hypnoid peristomes, but there are genera (for example, Bestia) with reduced peristomes. The calyptrae being frequently hairy is an unusual character in the flora area.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Costae short and double or rarely single in isolated leaves; stem and branch leaves differentiated.
Tripterocladium
1. Costae single; stem and branch leaves similar
→ 2
2. Medial laminal cells less than 4:1; capsules erect; peristome reduced.
Bestia
2. Medial laminal cells more than 5:1; capsules suberect, inclined, or rarely erect; peristome perfect.
Isothecium
Source FNA vol. 28, p. 620. FNA vol. 28, p. 616. Author: William R. Buck.
Parent taxa Lembophyllaceae > Isothecium
Sibling taxa
I. alopecuroides, I. cardotii, I. cristatum, I. stoloniferum
Subordinate taxa
Bestia, Isothecium, Tripterocladium
Synonyms Hypnum myosuroides, Pseudisothecium myosuroides
Name authority Bridel: Bryol. Univ. 2: 369. (1827) Brotherus
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