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Gulf Coast bulrush

bulrush, club-rush

Habit Plants annual; rhizomes absent. Herbs, annual or perennial, cespitose, rhizomatous or not, smooth, glabrous.
Culms

2–20 cm × 0.2–0.3 mm.

terete.

Leaves

sheaths green to stramineous;

distal blade mostly much longer than sheath, to 3 cm × 0.2 mm.

all basal;

sheaths green to stramineous, sometimes reddish proximally;

ligules absent;

blades rudimentary to exceeding culms.

Inflorescences

involucral bract 1, 3–10(–25) mm, rarely a second bract to 5 mm.

terminal, sometimes pseudolateral, capitate or solitary spikelet;

spikelets 1–3(–15);

involucral bracts 1(–2), spreading to erect, like foliage leaf blades.

Spikelets

2–8 × 1–1.5 mm;

scales colorless, stramineous, or pale greenish, markedly gibbous, veinless or obscurely 3–7-veined, midrib keeled, finely reticulate at 40X, membranous, hyaline, apex rounded, mucronate, or awned;

proximal scale to 1.5(–5) mm, awn to 3 mm;

other scales 1–1.2 mm, apex, mucro 0.1 mm.

scales 8–25, spirally arranged, each subtending flower.

Flowers

anthers 0.2 mm;

styles 3-fid.

bisexual;

perianth absent;

stamens 1–3;

styles linear, 2–3-fid, base persistent, sometimes slightly enlarged.

Achenes

often falling with and clasped by floral scales, whitish (or orange-brown when unripe), broadly ellipsoid to obovoid or outline subcircular, nearly equilaterally trigonous, faces convex, 0.7–0.9 × 0.5–0.6 mm, minutely papillose at 40X, papillae in many vertical rows or usually obscured by thick whitish surface layer.

biconvex or trigonous, papillose or longitudinally ribbed.

Isolepis pseudosetacea

Isolepis

Phenology Fruiting spring–summer.
Habitat Wet, freshwater, often drying, places in grasslands, open woods, limestone barrens, cultivated fields, waste places
Elevation 0(–100) m (0(–300) ft)
Distribution
from FNA
AL; AR; GA; LA; MO; MS; TX; s Europe; n Africa
[BONAP county map]
from USDA
Worldwide in cool-tropical and temperate regions; especially Africa and Australia
[BONAP county map]
Discussion

Isolepis pseudosetacea is often mistaken for the very similar I. carinata, with which it sometimes grows but does not intergrade. It may easily be distinguished by the characters given in the key.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 69 (4 in the flora).

Isolepis is difficult to delimit on a worldwide basis and has been included in Scirpus in the broad sense. Data derived from embryologic, genetic, and other studies led in recent years to the acceptance of Isolepis as a distinct genus (J. J. Bruhl 1995; P. Goetghebeur 1998; A. M. Muasya et al. 2001).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spikelets with scales markedly gibbous, often clasping shed achenes, colorless to stramineous, or orangish, or greenish; achenes acutely equilaterally trigonous; leaf sheaths green to stramineous or brown.
→ 2
1. Spikelets with scales neither gibbous nor clasping shed achenes, usually at least partly orange- to red-brown or blackish; achenes compressed-trigonous or thickly plano-convex; leaf sheaths usually reddish proximally.
→ 3
2. Scales from middle of spikelet 1.8–2 mm, with awns 0.2–0.5 mm; achenes 1–1.5 mm.
I. carinata
2. Scales from middle of spikelet 1–1.2 mm, with mucros to 0.1 mm; achenes 0.7–0.9 mm.
I. pseudosetacea
3. Achenes papillose, not ribbed; culms, leaves, and involucral bracts not orange-punctate or sheaths sparsely so.
I. cernua
3. Achenes not papillose, prominently longitudinally ribbed at 10X and minutely transversely ridged at 20–30X; culms, leaves, and involucral bracts orange-punctate at 10– 15X.
I. setacea
Source FNA vol. 23, p. 138. FNA vol. 23, p. 137. Author: S. Galen Smith.
Parent taxa Cyperaceae > Isolepis Cyperaceae
Sibling taxa
I. carinata, I. cernua, I. setacea
Subordinate taxa
I. carinata, I. cernua, I. pseudosetacea, I. setacea
Synonyms Scirpus pseudosetaceus, I. molesta, Scirpus molestus Scirpus section I.
Name authority (Daveau) Gandoger: Cat. Pl. Espagne, 331. (1917) R. Brown: Prodr., 221. (1810)
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