Isolepis |
Isolepis cernua |
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bulrush, club-rush |
fiberoptic grass, low bulrush, low clubrush, low lateral clubrush, tuft clubrush |
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Habit | Herbs, annual or perennial, cespitose, rhizomatous or not, smooth, glabrous. | Plants annual (or perennial?); rhizomes usually obscured by culm bases and very short, sometimes vertical and elongated. | ||||||||||||
Culms | terete. |
4–40 cm × 0.2–0.5 mm. |
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Leaves | all basal; sheaths green to stramineous, sometimes reddish proximally; ligules absent; blades rudimentary to exceeding culms. |
sometimes sparsely orange-punctate at 10–15X; sheaths usually reddish proximally; distal blade rudimentary to much longer than sheath, often exceeding culm, to 20 cm × 0.2–1 mm. |
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Inflorescences | terminal, sometimes pseudolateral, capitate or solitary spikelet; spikelets 1–3(–15); involucral bracts 1(–2), spreading to erect, like foliage leaf blades. |
involucral bract 1, sometimes subtending flower or resembling enlarged floral scale, 2–6(–23) mm. |
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Spikelets | scales 8–25, spirally arranged, each subtending flower. |
2–5 × 1–2 mm; scales partly or completely dark orange to red-brown, rarely stramineous, midrib greenish to stramineous, not gibbous, obscurely to prominently 3–11-veined, midrib keeled near apex, membranous, hyaline, apex rounded to acute, with mucro less than 0.1 mm; proximal scale to 2 mm; other scales 1.2–1.8 × 1–1.3 mm. |
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Flowers | bisexual; perianth absent; stamens 1–3; styles linear, 2–3-fid, base persistent, sometimes slightly enlarged. |
anthers 0.3–0.6 mm; styles 3-fid or 3-fid and 2-fid. |
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Achenes | biconvex or trigonous, papillose or longitudinally ribbed. |
falling separately from scales, medium to dark brown or stramineous, ellipsoid to obovoid, compressed-trigonous to thickly biconvex, lateral angles usually prominent, abaxial angle prominent to obscure, faces convex or adaxial face slightly concave, 0.8–1 × 0.5–0.7 mm, distinctly papillose at 10–15X to obscurely papillose at 40X, often with thin whitish surface layer. |
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2n | = 30. |
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Isolepis |
Isolepis cernua |
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Phenology | Fruiting late spring–winter (Pacific Coast), winter–spring (Texas). | |||||||||||||
Habitat | Wet, freshwater to brackish places on beaches, dunes, marine bluffs, sandy areas, mostly coastal | |||||||||||||
Elevation | 0–800 m (0–2600 ft) | |||||||||||||
Distribution |
Worldwide in cool-tropical and temperate regions; especially Africa and Australia |
CA; OR; TX; WA; BC; Mexico (Baja California); temperate South America; Eurasia; Africa; Australia; New Zealand
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Discussion | Species 69 (4 in the flora). Isolepis is difficult to delimit on a worldwide basis and has been included in Scirpus in the broad sense. Data derived from embryologic, genetic, and other studies led in recent years to the acceptance of Isolepis as a distinct genus (J. J. Bruhl 1995; P. Goetghebeur 1998; A. M. Muasya et al. 2001). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Isolepis cernua is widespread and variable. Four varieties were recognized by A. M. Muasya and D. M. Simpson (2002). Only var. ceruna is known from North America. The earliest collection I have seen from the Pacific Coast is from 1888; the earliest collection I have seen from Texas is from 1974. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 23, p. 137. | FNA vol. 23, p. 139. | ||||||||||||
Parent taxa | ||||||||||||||
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Synonyms | Scirpus section I. | Scirpus cernuus, Scirpus cernuus var. californicus, Scirpus cernuus subsp. californicus | ||||||||||||
Name authority | R. Brown: Prodr., 221. (1810) | (Vahl) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 106. (1817) | ||||||||||||
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