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bulrush, club-rush

fiberoptic grass, low bulrush, low clubrush, low lateral clubrush, tuft clubrush

Habit Herbs, annual or perennial, cespitose, rhizomatous or not, smooth, glabrous. Plants annual (or perennial?); rhizomes usually obscured by culm bases and very short, sometimes vertical and elongated.
Culms

terete.

4–40 cm × 0.2–0.5 mm.

Leaves

all basal;

sheaths green to stramineous, sometimes reddish proximally;

ligules absent;

blades rudimentary to exceeding culms.

sometimes sparsely orange-punctate at 10–15X;

sheaths usually reddish proximally;

distal blade rudimentary to much longer than sheath, often exceeding culm, to 20 cm × 0.2–1 mm.

Inflorescences

terminal, sometimes pseudolateral, capitate or solitary spikelet;

spikelets 1–3(–15);

involucral bracts 1(–2), spreading to erect, like foliage leaf blades.

involucral bract 1, sometimes subtending flower or resembling enlarged floral scale, 2–6(–23) mm.

Spikelets

scales 8–25, spirally arranged, each subtending flower.

2–5 × 1–2 mm;

scales partly or completely dark orange to red-brown, rarely stramineous, midrib greenish to stramineous, not gibbous, obscurely to prominently 3–11-veined, midrib keeled near apex, membranous, hyaline, apex rounded to acute, with mucro less than 0.1 mm;

proximal scale to 2 mm;

other scales 1.2–1.8 × 1–1.3 mm.

Flowers

bisexual;

perianth absent;

stamens 1–3;

styles linear, 2–3-fid, base persistent, sometimes slightly enlarged.

anthers 0.3–0.6 mm;

styles 3-fid or 3-fid and 2-fid.

Achenes

biconvex or trigonous, papillose or longitudinally ribbed.

falling separately from scales, medium to dark brown or stramineous, ellipsoid to obovoid, compressed-trigonous to thickly biconvex, lateral angles usually prominent, abaxial angle prominent to obscure, faces convex or adaxial face slightly concave, 0.8–1 × 0.5–0.7 mm, distinctly papillose at 10–15X to obscurely papillose at 40X, often with thin whitish surface layer.

2n

= 30.

Isolepis

Isolepis cernua

Phenology Fruiting late spring–winter (Pacific Coast), winter–spring (Texas).
Habitat Wet, freshwater to brackish places on beaches, dunes, marine bluffs, sandy areas, mostly coastal
Elevation 0–800 m (0–2600 ft)
Distribution
from USDA
Worldwide in cool-tropical and temperate regions; especially Africa and Australia
[BONAP county map]
from FNA
CA; OR; TX; WA; BC; Mexico (Baja California); temperate South America; Eurasia; Africa; Australia; New Zealand
[WildflowerSearch map]
[BONAP county map]
Discussion

Species 69 (4 in the flora).

Isolepis is difficult to delimit on a worldwide basis and has been included in Scirpus in the broad sense. Data derived from embryologic, genetic, and other studies led in recent years to the acceptance of Isolepis as a distinct genus (J. J. Bruhl 1995; P. Goetghebeur 1998; A. M. Muasya et al. 2001).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Isolepis cernua is widespread and variable. Four varieties were recognized by A. M. Muasya and D. M. Simpson (2002). Only var. ceruna is known from North America. The earliest collection I have seen from the Pacific Coast is from 1888; the earliest collection I have seen from Texas is from 1974.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spikelets with scales markedly gibbous, often clasping shed achenes, colorless to stramineous, or orangish, or greenish; achenes acutely equilaterally trigonous; leaf sheaths green to stramineous or brown.
→ 2
1. Spikelets with scales neither gibbous nor clasping shed achenes, usually at least partly orange- to red-brown or blackish; achenes compressed-trigonous or thickly plano-convex; leaf sheaths usually reddish proximally.
→ 3
2. Scales from middle of spikelet 1.8–2 mm, with awns 0.2–0.5 mm; achenes 1–1.5 mm.
I. carinata
2. Scales from middle of spikelet 1–1.2 mm, with mucros to 0.1 mm; achenes 0.7–0.9 mm.
I. pseudosetacea
3. Achenes papillose, not ribbed; culms, leaves, and involucral bracts not orange-punctate or sheaths sparsely so.
I. cernua
3. Achenes not papillose, prominently longitudinally ribbed at 10X and minutely transversely ridged at 20–30X; culms, leaves, and involucral bracts orange-punctate at 10– 15X.
I. setacea
Source FNA vol. 23, p. 137. Author: S. Galen Smith. FNA vol. 23, p. 139.
Parent taxa Cyperaceae Cyperaceae > Isolepis
Sibling taxa
I. carinata, I. pseudosetacea, I. setacea
Subordinate taxa
I. carinata, I. cernua, I. pseudosetacea, I. setacea
Synonyms Scirpus section I. Scirpus cernuus, Scirpus cernuus var. californicus, Scirpus cernuus subsp. californicus
Name authority R. Brown: Prodr., 221. (1810) (Vahl) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 106. (1817)
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