Isatis |
Isatis tinctoria |
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woad |
asp-of-Jerusalem, dyer's woad, woad |
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Habit | Biennials [annuals, perennials]; not scapose; (often glaucous), glabrous or pubescent. | Plants glaucous, usually glabrous, sometimes pubescent proximally. |
Stems | erect, often unbranched basally, paniculately branched distally. |
(3–)4–10(–15) dm. |
Leaves | basal and cauline; petiolate or sessile; basal rosulate [or not rosulate], petiolate [rarely sessile], blade margins entire, repand, or dentate [rarely pinnately lobed]; cauline blade (base auriculate, sagittate, [or amplexicaul, rarely attenuate]), margins entire [dentate]. |
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Basal leaves | petiole 0.5–5.5 cm; blade oblong or oblanceolate, (2.5–)5–15(–20) cm × (5–)15–35(–50) mm, base attenuate, margins entire, repand, or dentate, apex obtuse. |
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Cauline leaves | blade usually oblong or lanceolate, rarely linear-oblong, base sagittate or auriculate, apex acute. |
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Racemes | (corymbose, in panicles, several-flowered), considerably elongated in fruit. |
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Flowers | sepals erect or ascending, oblong [ovate]; petals oblanceolate [obovate, spatulate, or oblong], (equal to or longer than sepals), claw absent, (apex obtuse [subemarginate]); stamens slightly tetradynamous; filaments not dilated basally; anthers oblong [ovate], (apex obtuse or apiculate); nectar glands (6) confluent, or (4) lateral and median. |
sepals 1.5–2.8 × 1–1.5 mm, glabrous; petals 2.5–4 × 0.9–1.5 mm, base attenuate; filaments 1–2.5 mm; anthers 0.5–0.7 mm. |
Fruiting pedicels | reflexed, slender, (filiform, often thickened and clavate apically). |
5–10 mm. |
Fruits | siliques or silicles (samaroid), sessile, oblong, oblanceolate, elliptic, or obovate [ovate, cordate, spatulate, orbicular], 1- (or 2-)seeded, smooth, strongly angustiseptate, (prominently winged all around or distally; seed-bearing locule papery or corky, distinctly or obscurely 1–3-veined, sometimes keeled or shortly winged), glabrous or pubescent; valves and replum united; septum absent; ovules 1 (or 2) per ovary, (subapical); stigma capitate. |
black or dark brown, often broader distal to middle, (0.9–)1.1–2(–2.7) cm × 3–6(–10) mm, base cuneate, margins sometimes slightly constricted, apex usually subacute or rounded, rarely subemarginate; locule with distinct midvein, lateral veins inconspicuous, 3–6(–10) mm; apical wing 3.5–5(–7) mm wide. |
Seeds | plump, not winged, narrowly oblong; seed coat (smooth), not mucilaginous when wetted; cotyledons incumbent or accumbent. |
light brown, 2.3–3.5(–4.5) × 0.8–1 mm. |
2n | = 14, 28. |
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Isatis |
Isatis tinctoria |
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Phenology | Flowering Apr–Jun. | |
Habitat | Roadsides, fields, pastures, sagebrush hillsides, prairies, railroad embankments, waste places | |
Elevation | 300-2200 m (1000-7200 ft) | |
Distribution |
Europe; c Asia; sw Asia; n Africa [Introduced in North America; introduced also in South America (Chile, Peru)] |
CA; ID; IL; MO; MT; NM; NV; NY; OR; UT; VA; WA; WV; WY; BC; NF; ON; QC; Europe; c Asia; sw Asia; n Africa [Introduced in North America; introduced also in South America (Chile, Peru)]
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Discussion | Species 50 (1 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Isatis tinctoria has been cultivated since ancient times as a source of a blue dye (woad) obtained by fermenting the ground leaves and proximal portions of the plant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 567. | FNA vol. 7, p. 568. |
Parent taxa | ||
Subordinate taxa | ||
Name authority | Linnaeus: Sp. Pl. 2: 670. (1753): Gen. Pl. ed. 5, 301. (1754) | Linnaeus: Sp. Pl. 2: 670. (1753) |
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