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woad

asp-of-Jerusalem, dyer's woad, woad

Habit Biennials [annuals, perennials]; not scapose; (often glaucous), glabrous or pubescent. Plants glaucous, usually glabrous, sometimes pubescent proximally.
Stems

erect, often unbranched basally, paniculately branched distally.

(3–)4–10(–15) dm.

Leaves

basal and cauline;

petiolate or sessile;

basal rosulate [or not rosulate], petiolate [rarely sessile], blade margins entire, repand, or dentate [rarely pinnately lobed];

cauline blade (base auriculate, sagittate, [or amplexicaul, rarely attenuate]), margins entire [dentate].

Basal leaves

petiole 0.5–5.5 cm;

blade oblong or oblanceolate, (2.5–)5–15(–20) cm × (5–)15–35(–50) mm, base attenuate, margins entire, repand, or dentate, apex obtuse.

Cauline leaves

blade usually oblong or lanceolate, rarely linear-oblong, base sagittate or auriculate, apex acute.

Racemes

(corymbose, in panicles, several-flowered), considerably elongated in fruit.

Flowers

sepals erect or ascending, oblong [ovate];

petals oblanceolate [obovate, spatulate, or oblong], (equal to or longer than sepals), claw absent, (apex obtuse [subemarginate]);

stamens slightly tetradynamous;

filaments not dilated basally;

anthers oblong [ovate], (apex obtuse or apiculate);

nectar glands (6) confluent, or (4) lateral and median.

sepals 1.5–2.8 × 1–1.5 mm, glabrous;

petals 2.5–4 × 0.9–1.5 mm, base attenuate;

filaments 1–2.5 mm;

anthers 0.5–0.7 mm.

Fruiting pedicels

reflexed, slender, (filiform, often thickened and clavate apically).

5–10 mm.

Fruits

siliques or silicles (samaroid), sessile, oblong, oblanceolate, elliptic, or obovate [ovate, cordate, spatulate, orbicular], 1- (or 2-)seeded, smooth, strongly angustiseptate, (prominently winged all around or distally; seed-bearing locule papery or corky, distinctly or obscurely 1–3-veined, sometimes keeled or shortly winged), glabrous or pubescent;

valves and replum united;

septum absent;

ovules 1 (or 2) per ovary, (subapical);

stigma capitate.

black or dark brown, often broader distal to middle, (0.9–)1.1–2(–2.7) cm × 3–6(–10) mm, base cuneate, margins sometimes slightly constricted, apex usually subacute or rounded, rarely subemarginate;

locule with distinct midvein, lateral veins inconspicuous, 3–6(–10) mm;

apical wing 3.5–5(–7) mm wide.

Seeds

plump, not winged, narrowly oblong;

seed coat (smooth), not mucilaginous when wetted;

cotyledons incumbent or accumbent.

light brown, 2.3–3.5(–4.5) × 0.8–1 mm.

2n

= 14, 28.

Isatis

Isatis tinctoria

Phenology Flowering Apr–Jun.
Habitat Roadsides, fields, pastures, sagebrush hillsides, prairies, railroad embankments, waste places
Elevation 300-2200 m (1000-7200 ft)
Distribution
from USDA
Europe; c Asia; sw Asia; n Africa [Introduced in North America; introduced also in South America (Chile, Peru)]
[BONAP county map]
from FNA
CA; ID; IL; MO; MT; NM; NV; NY; OR; UT; VA; WA; WV; WY; BC; NF; ON; QC; Europe; c Asia; sw Asia; n Africa [Introduced in North America; introduced also in South America (Chile, Peru)]
[WildflowerSearch map]
[BONAP county map]
Discussion

Species 50 (1 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Isatis tinctoria has been cultivated since ancient times as a source of a blue dye (woad) obtained by fermenting the ground leaves and proximal portions of the plant.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 567. Author: Ihsan A. Al-Shehbaz. FNA vol. 7, p. 568.
Parent taxa Brassicaceae > tribe Isatideae Brassicaceae > tribe Isatideae > Isatis
Subordinate taxa
I. tinctoria
Name authority Linnaeus: Sp. Pl. 2: 670. (1753): Gen. Pl. ed. 5, 301. (1754) Linnaeus: Sp. Pl. 2: 670. (1753)
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