Isatis |
Brassicaceae tribe Isatideae |
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woad |
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Habit | Biennials [annuals, perennials]; not scapose; (often glaucous), glabrous or pubescent. | Annuals or biennials [perennials]; eglandular. |
Stems | erect, often unbranched basally, paniculately branched distally. |
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Leaves | basal and cauline; petiolate or sessile; basal rosulate [or not rosulate], petiolate [rarely sessile], blade margins entire, repand, or dentate [rarely pinnately lobed]; cauline blade (base auriculate, sagittate, [or amplexicaul, rarely attenuate]), margins entire [dentate]. |
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Cauline leaves | sessile [petiolate]; blade base auriculate [not auriculate], margins usually entire [dentate]. |
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Trichomes | simple or absent. |
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Racemes | (corymbose, in panicles, several-flowered), considerably elongated in fruit. |
ebracteate, elongated in fruit. |
Flowers | sepals erect or ascending, oblong [ovate]; petals oblanceolate [obovate, spatulate, or oblong], (equal to or longer than sepals), claw absent, (apex obtuse [subemarginate]); stamens slightly tetradynamous; filaments not dilated basally; anthers oblong [ovate], (apex obtuse or apiculate); nectar glands (6) confluent, or (4) lateral and median. |
actinomorphic; sepals erect or ascending [spreading], lateral pair not saccate or subsaccate basally; petals yellow [white], claw usually present, rarely absent, obscure; filaments unappendaged, not winged; pollen 3-colpate. |
Fruiting pedicels | reflexed, slender, (filiform, often thickened and clavate apically). |
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Fruits | siliques or silicles (samaroid), sessile, oblong, oblanceolate, elliptic, or obovate [ovate, cordate, spatulate, orbicular], 1- (or 2-)seeded, smooth, strongly angustiseptate, (prominently winged all around or distally; seed-bearing locule papery or corky, distinctly or obscurely 1–3-veined, sometimes keeled or shortly winged), glabrous or pubescent; valves and replum united; septum absent; ovules 1 (or 2) per ovary, (subapical); stigma capitate. |
siliques or silicles (samaroid), indehiscent, unsegmented, angustiseptate, (woody); ovules 1 or 2 per ovary; style usually absent, rarely distinct; stigma entire. |
Seeds | plump, not winged, narrowly oblong; seed coat (smooth), not mucilaginous when wetted; cotyledons incumbent or accumbent. |
aseriate; cotyledons accumbent or incumbent. |
Isatis |
Brassicaceae tribe Isatideae |
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Distribution |
Europe; c Asia; sw Asia; n Africa [Introduced in North America; introduced also in South America (Chile, Peru)] |
Europe; Asia; n Africa [Introduced in North America] |
Discussion | Species 50 (1 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 2–4, species 90–95 (2 genera, 2 species in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 567. | FNA vol. 7, p. 567. |
Parent taxa | ||
Subordinate taxa | ||
Name authority | Linnaeus: Sp. Pl. 2: 670. (1753): Gen. Pl. ed. 5, 301. (1754) | de Candolle: Mém. Mus. Hist. Nat. 7: 241. (1821) |
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