Iridaceae |
Romulea |
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iris family |
romulea, sand-crocus |
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Habit | Herbs, perennial, rarely annual [or shrubs with woody caudex], evergreen or seasonal, sometimes cespitose; rootstock a rhizome, bulb, or corm. | Herbs, perennial, from corms. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | true stem aerial [subterranean and only peduncles of individual flowers aerial], simple or branched. |
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Flowering stems | aerial (or subterranean in Romulea), simple or branched, terete or variously compressed, angled or winged. |
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Leaves | basal and cauline, distichous; proximal 2–3 sometimes membranous, not reaching much above ground; others with open or closed sheaths, usually unifacial [bifacial or terete], oriented edgewise to the stem; blade parallel-veined, plane or pleated, channeled. |
1–several; blade linear, oval to terete in cross section, 4-grooved (2 on each surface, on either side of thickened midrib) [2-grooved]. |
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Inflorescences | umbellate, monochasial cymes (rhipidia), spikes, or solitary flowers; rhipidia enclosed in 2, opposed, usually large, leafy to dry bracts (spathes); flowers except for the first subtended by 1 floral bract; spike flowers each subtended by 2, opposed bracts. |
solitary flowers, interpreted as sessile, thus without pedicels; bracts inserted at ovary base, green or flushed with purple, unequal, outer exceeding inner, firm, inner bract margins membranous to scarious (rarely entirely scarious). |
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Flowers | usually pedicellate [± sessile]; perianth actinomorphic or zygomorphic, petaloid, with 2 equal or unequal whorls of 3 tepals each [1 whorl of 6]; tepals usually large, showy, distinct or connate in tube; stamens 3 [2], inserted at base of outer tepals or in tube, symmetrically arranged or unilateral, arcuate [declinate]; filaments distinct or partly to completely connate, sometimes weak, unable to support anthers; anthers with 2 pollen sacs, extrorse, occasionally latrorse, usually dehiscing longitudinally [rarely apically]; ovary inferior [superior in Tasmanian Isophysis], 3-locular [1-locular]; placentation axile [parietal]; ovules 2–few, anatropous; style single, filiform at least proximally, usually 3-branched or 3-lobed, branches either filiform, distally expanded, sometimes each divided in distal 1/2, stigmatic toward apices, or branches thickened, or flattened, petaloid, stigmas then abaxial below apices. |
odorless [fragrant], actinomorphic; tepals forming wide cup, connate into tube, subequal, outer whorl often slightly larger than inner; perianth tube funnel-shaped [or cylindric]; stamens symmetrical; filaments distinct [connate]; anthers usually erect, contiguous; style branching opposite or beyond anthers into 3 slender branches divided for ± 1/2 length. |
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Fruits | capsular, loculicidal, rarely indehiscent, firm to cartilaginous, occasionally woody. |
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Capsules | ovoid-oblong, walls firm, cartilaginous. |
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Seeds | globose to angular (prismatic) or discoid, sometimes broadly winged; seed coat usually dry (rarely fleshy); endosperm hard, with reserves of hemicellulose, oil, and protein; embryo small. |
many, globose; seed coat light to dark brown, often smooth, shiny. |
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x | = 13 or 14. |
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Iridaceae |
Romulea |
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Distribution | Nearly worldwide but rare in tropical lowlands; best represented in southern Africa |
Africa; s Europe; Middle East [Introduced in North America] |
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Discussion | Genera ca. 65, species ca. 1810 (16 genera, 92 species; 9 genera, 19 species introduced, and various hybrid complexes in the flora). Iridaceae are currently divided into four subfamilies (P. Goldblatt 1990, 1991). Subfamily Isophysidoideae Takhtajan is monotypic, comprising the Tasmanian Isophysis T. Moore with a superior ovary. Only subfamily Iridoideae is native in North America. The remaining Nivenioideae Schulze ex Goldblatt and Ixioideae Klatt are centered in Africa south of the Sahara. Iridaceae are of considerable economic importance in ornamental horticulture and the cut-flower industry, especially Iris, Gladiolus, and Freesia. Several other genera (e.g., Crocus, Dietes, Sparaxis, Tritonia, Watsonia) are cultivated in gardens in both tropical and temperate areas. Moraea and Homeria are poisonous and pose significant problems in cattle- and sheep-raising areas, notably in southern Africa. In addition to the several genera and species escaped from cultivation and dealt with in detail below, the following are widely grown in areas of mild winter and may persist in and near abandoned gardens, sometimes reproducing successfully: Dietes Salisbury [D. iridioides (Linnaeus) Salisbury ex Klatt, D. grandiflora N. E. Brown]; Ixia Linnaeus (I. maculata Linnaeus, I. polystachya Linnaeus); Crocus cultivars and even some wild species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 90 (1 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 26, p. 348. | FNA vol. 26, p. 407. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Name authority | Jussieu | Maratti: Pl. Romul. Saturn., 13. (1772) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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