Iridaceae |
Gladiolus |
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iris family |
corn-flag, gladiolus |
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Habit | Herbs, perennial, rarely annual [or shrubs with woody caudex], evergreen or seasonal, sometimes cespitose; rootstock a rhizome, bulb, or corm. | Herbs, perennial, from corms. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | simple or branched. |
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Flowering stems | aerial (or subterranean in Romulea), simple or branched, terete or variously compressed, angled or winged. |
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Leaves | basal and cauline, distichous; proximal 2–3 sometimes membranous, not reaching much above ground; others with open or closed sheaths, usually unifacial [bifacial or terete], oriented edgewise to the stem; blade parallel-veined, plane or pleated, channeled. |
1–9; blade lanceolate to linear, plane or margins and/or midribs variously raised and thickened (then H- or X-shaped in cross section), or evidently terete, midribs and margins much thickened, grooved; grooves 4, narrow, longitudinal. |
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Inflorescences | umbellate, monochasial cymes (rhipidia), spikes, or solitary flowers; rhipidia enclosed in 2, opposed, usually large, leafy to dry bracts (spathes); flowers except for the first subtended by 1 floral bract; spike flowers each subtended by 2, opposed bracts. |
spicate, partly to fully secund or with flowers weakly distichous; bracts green, sometimes flushed grayish purple, unequal, outer usually exceeding inner, acute or inner forked or notched apically. |
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Flowers | usually pedicellate [± sessile]; perianth actinomorphic or zygomorphic, petaloid, with 2 equal or unequal whorls of 3 tepals each [1 whorl of 6]; tepals usually large, showy, distinct or connate in tube; stamens 3 [2], inserted at base of outer tepals or in tube, symmetrically arranged or unilateral, arcuate [declinate]; filaments distinct or partly to completely connate, sometimes weak, unable to support anthers; anthers with 2 pollen sacs, extrorse, occasionally latrorse, usually dehiscing longitudinally [rarely apically]; ovary inferior [superior in Tasmanian Isophysis], 3-locular [1-locular]; placentation axile [parietal]; ovules 2–few, anatropous; style single, filiform at least proximally, usually 3-branched or 3-lobed, branches either filiform, distally expanded, sometimes each divided in distal 1/2, stigmatic toward apices, or branches thickened, or flattened, petaloid, stigmas then abaxial below apices. |
somewhat fragrant, zygomorphic [actinomorphic]; tepals basally connate into tube, variously colored, usually with contrasting markings comprising nectar guide on outer tepals, usually unequal, dorsal tepal largest, arched to hooded over stamens, outer 3 tepals narrower; perianth tube obliquely funnel-shaped to cylindric; stamens usually unilateral; anthers usually parallel; style usually arching over stamens, dividing into 3 filiform branches, these distally expanded. |
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Fruits | capsular, loculicidal, rarely indehiscent, firm to cartilaginous, occasionally woody. |
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Capsules | usually slightly inflated, oblong to ellipsoid or globose [rarely nearly cylindric], softly cartilaginous. |
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Seeds | globose to angular (prismatic) or discoid, sometimes broadly winged; seed coat usually dry (rarely fleshy); endosperm hard, with reserves of hemicellulose, oil, and protein; embryo small. |
usually many, broadly winged; rarely few, wingless, globose or angular; seed coat light to dark brown. |
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x | = 15. |
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Iridaceae |
Gladiolus |
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Distribution | Nearly worldwide but rare in tropical lowlands; best represented in southern Africa |
Africa; Madagascar; Eurasia [Introduced in North America] |
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Discussion | Genera ca. 65, species ca. 1810 (16 genera, 92 species; 9 genera, 19 species introduced, and various hybrid complexes in the flora). Iridaceae are currently divided into four subfamilies (P. Goldblatt 1990, 1991). Subfamily Isophysidoideae Takhtajan is monotypic, comprising the Tasmanian Isophysis T. Moore with a superior ovary. Only subfamily Iridoideae is native in North America. The remaining Nivenioideae Schulze ex Goldblatt and Ixioideae Klatt are centered in Africa south of the Sahara. Iridaceae are of considerable economic importance in ornamental horticulture and the cut-flower industry, especially Iris, Gladiolus, and Freesia. Several other genera (e.g., Crocus, Dietes, Sparaxis, Tritonia, Watsonia) are cultivated in gardens in both tropical and temperate areas. Moraea and Homeria are poisonous and pose significant problems in cattle- and sheep-raising areas, notably in southern Africa. In addition to the several genera and species escaped from cultivation and dealt with in detail below, the following are widely grown in areas of mild winter and may persist in and near abandoned gardens, sometimes reproducing successfully: Dietes Salisbury [D. iridioides (Linnaeus) Salisbury ex Klatt, D. grandiflora N. E. Brown]; Ixia Linnaeus (I. maculata Linnaeus, I. polystachya Linnaeus); Crocus cultivars and even some wild species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 260 (4 in the flora). Hybrid strains of Gladiolus are widely cultivated, and some wild species are grown outdoors in areas with mild winters. The common, large-flowered hybrids are the product of crossing between four or five species, followed by selection. Named variously G. gandavensis L. van Houtte, G. lemoinei Baker, and G. hortulanus L. H. Bailey, these plants are occasionally found in and near gardens and cemeteries and are easily recognized by their large, brightly colored flowers with the inner tepals 60–70 mm long and the outer tepals about 50 mm long. They reproduce mainly vegetatively and persist for some years where winters are mild; they show no evidence of spreading into native vegetation. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 26, p. 348. | FNA vol. 26, p. 407. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Name authority | Jussieu | Linnaeus: Sp. Pl. 1: 36. (1753): Gen. Pl. ed. 5, 23. (1754) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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