Iridaceae |
Freesia |
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iris family |
freesia |
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Habit | Herbs, perennial, rarely annual [or shrubs with woody caudex], evergreen or seasonal, sometimes cespitose; rootstock a rhizome, bulb, or corm. | Herbs, perennial, from corms. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | simple or branched. |
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Flowering stems | aerial (or subterranean in Romulea), simple or branched, terete or variously compressed, angled or winged. |
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Leaves | basal and cauline, distichous; proximal 2–3 sometimes membranous, not reaching much above ground; others with open or closed sheaths, usually unifacial [bifacial or terete], oriented edgewise to the stem; blade parallel-veined, plane or pleated, channeled. |
several, sometimes prostrate; blade plane, lanceolate [ensiform or oblong]. |
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Inflorescences | umbellate, monochasial cymes (rhipidia), spikes, or solitary flowers; rhipidia enclosed in 2, opposed, usually large, leafy to dry bracts (spathes); flowers except for the first subtended by 1 floral bract; spike flowers each subtended by 2, opposed bracts. |
spicate, secund, usually strongly flexed basally, ± horizontal, several–many-flowered; bract green [pale straw colored], coriaceous or membranous. |
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Flowers | usually pedicellate [± sessile]; perianth actinomorphic or zygomorphic, petaloid, with 2 equal or unequal whorls of 3 tepals each [1 whorl of 6]; tepals usually large, showy, distinct or connate in tube; stamens 3 [2], inserted at base of outer tepals or in tube, symmetrically arranged or unilateral, arcuate [declinate]; filaments distinct or partly to completely connate, sometimes weak, unable to support anthers; anthers with 2 pollen sacs, extrorse, occasionally latrorse, usually dehiscing longitudinally [rarely apically]; ovary inferior [superior in Tasmanian Isophysis], 3-locular [1-locular]; placentation axile [parietal]; ovules 2–few, anatropous; style single, filiform at least proximally, usually 3-branched or 3-lobed, branches either filiform, distally expanded, sometimes each divided in distal 1/2, stigmatic toward apices, or branches thickened, or flattened, petaloid, stigmas then abaxial below apices. |
often strongly fragrant, zygomorphic; tepals connate into tube, mostly white or yellow, sometimes pink or red, subequal, outer whorl slightly larger than inner; perianth tube cylindric or funnel-shaped, short or long proximally, abruptly expanded into short to long tube or flared distally; stamens asymmetrically disposed, unilateral; anthers usually parallel; style arching over filaments, dividing into 3 filiform branches. |
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Fruits | capsular, loculicidal, rarely indehiscent, firm to cartilaginous, occasionally woody. |
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Capsules | irregularly globose, usually rugose. |
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Seeds | globose to angular (prismatic) or discoid, sometimes broadly winged; seed coat usually dry (rarely fleshy); endosperm hard, with reserves of hemicellulose, oil, and protein; embryo small. |
several per locule, globose, without wings; seed coat light to dark brown, hard, shiny. |
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Corms | tunicate, conic, ca. 10 mm diam.; tunics fibrous; fibers fine, pale, reticulate. |
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x | = 11. |
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Iridaceae |
Freesia |
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Distribution | Nearly worldwide but rare in tropical lowlands; best represented in southern Africa |
South Africa [Introduced in North America] |
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Discussion | Genera ca. 65, species ca. 1810 (16 genera, 92 species; 9 genera, 19 species introduced, and various hybrid complexes in the flora). Iridaceae are currently divided into four subfamilies (P. Goldblatt 1990, 1991). Subfamily Isophysidoideae Takhtajan is monotypic, comprising the Tasmanian Isophysis T. Moore with a superior ovary. Only subfamily Iridoideae is native in North America. The remaining Nivenioideae Schulze ex Goldblatt and Ixioideae Klatt are centered in Africa south of the Sahara. Iridaceae are of considerable economic importance in ornamental horticulture and the cut-flower industry, especially Iris, Gladiolus, and Freesia. Several other genera (e.g., Crocus, Dietes, Sparaxis, Tritonia, Watsonia) are cultivated in gardens in both tropical and temperate areas. Moraea and Homeria are poisonous and pose significant problems in cattle- and sheep-raising areas, notably in southern Africa. In addition to the several genera and species escaped from cultivation and dealt with in detail below, the following are widely grown in areas of mild winter and may persist in and near abandoned gardens, sometimes reproducing successfully: Dietes Salisbury [D. iridioides (Linnaeus) Salisbury ex Klatt, D. grandiflora N. E. Brown]; Ixia Linnaeus (I. maculata Linnaeus, I. polystachya Linnaeus); Crocus cultivars and even some wild species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 15 (2 in the flora). Critical phylogenetic study has shown Freesia to be nested in Anomatheca, and the two genera have been united. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 26, p. 348. | FNA vol. 26, p. 405. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Anomatheca | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | Jussieu | Ecklon ex Klatt: Linnaea 34: 672. (1866) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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