Iridaceae |
Calydorea |
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iris family |
violet-lily |
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Habit | Herbs, perennial, rarely annual [or shrubs with woody caudex], evergreen or seasonal, sometimes cespitose; rootstock a rhizome, bulb, or corm. | Herbs, perennial, from tunicate, ovoid bulbs; tunic brown, dry, brittle, papery. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | simple or branched. |
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Flowering stems | aerial (or subterranean in Romulea), simple or branched, terete or variously compressed, angled or winged. |
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Leaves | basal and cauline, distichous; proximal 2–3 sometimes membranous, not reaching much above ground; others with open or closed sheaths, usually unifacial [bifacial or terete], oriented edgewise to the stem; blade parallel-veined, plane or pleated, channeled. |
few, basal larger; blade pleated, linear-lanceolate. |
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Inflorescences | umbellate, monochasial cymes (rhipidia), spikes, or solitary flowers; rhipidia enclosed in 2, opposed, usually large, leafy to dry bracts (spathes); flowers except for the first subtended by 1 floral bract; spike flowers each subtended by 2, opposed bracts. |
rhipidiate, few-flowered; spathes green, unequal, outer shorter than inner, apex usually brown, acute, dry. |
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Flowers | usually pedicellate [± sessile]; perianth actinomorphic or zygomorphic, petaloid, with 2 equal or unequal whorls of 3 tepals each [1 whorl of 6]; tepals usually large, showy, distinct or connate in tube; stamens 3 [2], inserted at base of outer tepals or in tube, symmetrically arranged or unilateral, arcuate [declinate]; filaments distinct or partly to completely connate, sometimes weak, unable to support anthers; anthers with 2 pollen sacs, extrorse, occasionally latrorse, usually dehiscing longitudinally [rarely apically]; ovary inferior [superior in Tasmanian Isophysis], 3-locular [1-locular]; placentation axile [parietal]; ovules 2–few, anatropous; style single, filiform at least proximally, usually 3-branched or 3-lobed, branches either filiform, distally expanded, sometimes each divided in distal 1/2, stigmatic toward apices, or branches thickened, or flattened, petaloid, stigmas then abaxial below apices. |
short-lived, erect, actinomorphic; tepals spreading from base, distinct, blue to mauve, ± equal [outer whorl considerably larger than inner]; stamens distinct or filaments variously connate; anthers sometimes connate basally; style eccentric [central] when flower fully open, recurving, slender, branching between middle of filaments and anther apices [branching distal to anthers or 3-lobed apically]; branches [lobes] ascending, undivided, short, stigmatic apically. |
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Fruits | capsular, loculicidal, rarely indehiscent, firm to cartilaginous, occasionally woody. |
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Capsules | ovoid to oblong, cartilaginous, apex truncate. |
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Seeds | globose to angular (prismatic) or discoid, sometimes broadly winged; seed coat usually dry (rarely fleshy); endosperm hard, with reserves of hemicellulose, oil, and protein; embryo small. |
many, prismatic; seed coat brown. |
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x | = 7. |
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Iridaceae |
Calydorea |
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Distribution | Nearly worldwide but rare in tropical lowlands; best represented in southern Africa |
South America; North America (incl Mexico) |
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Discussion | Genera ca. 65, species ca. 1810 (16 genera, 92 species; 9 genera, 19 species introduced, and various hybrid complexes in the flora). Iridaceae are currently divided into four subfamilies (P. Goldblatt 1990, 1991). Subfamily Isophysidoideae Takhtajan is monotypic, comprising the Tasmanian Isophysis T. Moore with a superior ovary. Only subfamily Iridoideae is native in North America. The remaining Nivenioideae Schulze ex Goldblatt and Ixioideae Klatt are centered in Africa south of the Sahara. Iridaceae are of considerable economic importance in ornamental horticulture and the cut-flower industry, especially Iris, Gladiolus, and Freesia. Several other genera (e.g., Crocus, Dietes, Sparaxis, Tritonia, Watsonia) are cultivated in gardens in both tropical and temperate areas. Moraea and Homeria are poisonous and pose significant problems in cattle- and sheep-raising areas, notably in southern Africa. In addition to the several genera and species escaped from cultivation and dealt with in detail below, the following are widely grown in areas of mild winter and may persist in and near abandoned gardens, sometimes reproducing successfully: Dietes Salisbury [D. iridioides (Linnaeus) Salisbury ex Klatt, D. grandiflora N. E. Brown]; Ixia Linnaeus (I. maculata Linnaeus, I. polystachya Linnaeus); Crocus cultivars and even some wild species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 12 (1 in the flora). The generic position of the single North American species of Calydorea has long been unsettled. Its segregation in the monotypic Salpingostylis (J. K. Small 1931b) is unsatisfactory. This Florida species differs little from those of Mexico, which in turn show strong similarities with others in South America. Some of these species were included by J. G. Baker (1892) in Sphenostigma, but the type of that genus was shown to be a species of Gelasine by P. Ravenna (1977), who then placed the Floridian and Mexican species that had been referred to Sphenostigma in Cardiostigma. P. Goldblatt and J. E. Henrich (1991) preferred to include all these species in a slightly expanded Calydorea, other species of which have the style dividing near the anther bases and the style branches somewhat longer than in the Mexican and Floridian species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 26, p. 348. | FNA vol. 26, p. 400. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Cardiostigma, Catila, Salpingostylis, Sphenostigma | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | Jussieu | Herbert: Edwards’s Bot. Reg. 29(misc.): 85. (1843) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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