The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

cypress-leaf claw-moss, cypress-leaf plaitmoss, hypnum moss

curly claw-moss, hypnum moss

Habit Plants small to large, rusty green, golden green, yellow-green, or pale green. Plants medium-sized, not in tufts, pale yellow-green to golden green.
Stem(s)

1–8+ cm, pale to yellowish green, brown with age, creeping to erect, complanate-foliate or not, irregularly pinnate to nearly unbranched, attached shoots often regularly pinnate, branches 1–3 cm;

hyalodermis absent, central strand poorly developed;

pseudoparaphyllia filamentous, 1–3-seriate at base.

leaves circinate-secund, ovate-lanceolate, gradually tapering to insertion, gradually narrowed to apex, 1.5–2.8 × 0.4–0.6 mm;

base not decurrent;

margins plane, sinuate proximally, serrate toward apex;

apex narrowly acuminate;

costa double, short, indistinct, less than 1/3 leaf length;

alar region slightly differentiated, of 3 or 4 cells at extreme basal angles;

basal laminal cells shorter, broader than medial cells, not strongly pigmented, walls pitted;

medial cells 50–80(–90) × 3–4 µm. Sexual condition dioicous;

inner perichaetial leaves lanceolate, weakly or not plicate, margins slightly toothed or entire near apex.

Leaves

strongly imbricate, not to decidedly falcate-secund, ovate to oblong-lanceolate, gradually or abruptly narrowed to apex, 1.5–2 × 0.5–0.8 mm;

base not decurrent, not auriculate;

margins recurved to plane proximally, serrulate (sometimes weakly) distally, occasionally nearly entire;

acumen slender;

costa double and short or obscure;

alar region well defined, basalmost cells larger, sometimes hyaline, yellowish or brownish;

basal laminal cells shorter, wider than medial cells, not pigmented, walls not pitted;

medial cells (50–)60–80 × 3–4(–5) µm. Sexual condition dioicous;

inner perichaetial leaves oblong-lanceolate, margins serrulate distally, costa obscure.

Seta

reddish, 1–2.5(–3) cm.

red-brown, 2.5–3 cm.

Capsule

slightly inclined, reddish, cylindric, 1.8–2.5(–2.8) mm;

annulus 1–3-seriate;

operculum conic to rostrate;

endostome cilia 1–2(–3).

erect to suberect, light brown, cylindric, 3–5 mm;

annulus 1–3-seriate;

operculum conic or rostellate;

endostome cilia unknown.

Hypnum cupressiforme

Hypnum subimponens

Phenology Capsules mature May–Jun.
Habitat Epiphytic on living trees, cliffs, vertical rock faces, decaying logs, terrestrial
Elevation low to moderate elevations (0-1500 m) (low to moderate elevations (0-4900 ft))
Distribution
Nearly worldwide; except Antarctica
[WildflowerSearch map]
from FNA
AK; CA; ID; MT; OR; WA; BC; NT; NU; YT; e Asia
[WildflowerSearch map]
Discussion

Varieties 9 (4 in the flora).

Hypnum cupressiforme is an extremely polymorphic species, reflected in the more than 60 varieties that have been described. The species has a wide ecological amplitude as well as a nearly cosmopolitan distribution and is found in all climatic regions except the Antarctic. Taxonomic features reliable in most other species of Hypnum are plastic in H. cupressiforme. Within a single clone, it is possible to sort out several named varieties. Variety lacunosum Bridel was noted by H. Ando (1989) to be weakly differentiated in North America, and he tentatively cited specimens from Nova Scotia, Newfoundland, and South Dakota. Ando noted that var. lacunosum typically is robust, thick-complanate to julaceous, with leaves almost straight to weakly falcate and abruptly narrowed to a short acumen, but the North American material is not robust and is therefore problematic.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Specimens of Hypnum subimponens from Arctic regions are problematic; in fact, any specimens from horizontal surfaces can be difficult to place accurately. Sporophytes, produced in spring, are frequent only on vertical surfaces and then often abundant. Unlike similar species in western North America, H. subimponens produces cylindric, erect capsules (slightly curved below the mouth when mature). Hypnum lindbergii has falcate-secund leaves, while those in H. subimponens are circinate-secund to strongly hamate. Key traits of the stem leaves separate H. subimponens from H. hamulosum, and the former is usually more than twice the size of the latter. See also discussions of 3. H. callichroum, 11. H. holmenii, and 16. H. plicatulum. Plants of H. subimponens have branches 0.5–1 mm wide; the alar cells are slightly bulging, hyaline, and thin-walled.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Branch leaves 1.5-2 × 0.4-0.6 mm or slightly larger
→ 2
1. Branch leaves 1-1.4 × 0.2-0.4 mm or slightly smaller
→ 3
2. Stems weakly complanate-foliate; leaves usually falcate, sometimes strongly so; alar cells not excavate or pigmented.
var. cupressiforme
2. Stems subjulaceous to complanate-foliate; leaves straight to weakly falcate; alar cells excavate, pigmented.
var. subjulaceum
3. Leaves usually falcate, sometimes strongly so; margins subentire to distinctly toothed distally.
var. cupressiforme
3. Leaves straight to somewhat falcate; margins entire or subentire
→ 4
4. Stems subjulaceous, creeping; leaves oblong-lanceolate, gradually narrowed to apex; laminal cells 60-80 × 3-4 µm (or slightly larger).
var. filiforme
4. Stems julaceous, creeping to erect; leaves ovate or widely oblong, abruptly narrowed to apex; laminal cells 40-50 × 5-6 µm.
var. julaceum
Source FNA vol. 28, p. 537. FNA vol. 28, p. 548.
Parent taxa Hypnaceae > Hypnum Hypnaceae > Hypnum
Sibling taxa
H. andoi, H. bambergeri, H. callichroum, H. circinale, H. curvifolium, H. dieckei, H. fauriei, H. fujiyamae, H. hamulosum, H. holmenii, H. imponens, H. jutlandicum, H. lindbergii, H. pallescens, H. plicatulum, H. pratense, H. procerrimum, H. recurvatum, H. revolutum, H. subimponens, H. vaucheri
H. andoi, H. bambergeri, H. callichroum, H. circinale, H. cupressiforme, H. curvifolium, H. dieckei, H. fauriei, H. fujiyamae, H. hamulosum, H. holmenii, H. imponens, H. jutlandicum, H. lindbergii, H. pallescens, H. plicatulum, H. pratense, H. procerrimum, H. recurvatum, H. revolutum, H. vaucheri
Subordinate taxa
H. cupressiforme var. cupressiforme, H. cupressiforme var. filiforme, H. cupressiforme var. julaceum, H. cupressiforme var. subjulaceum
Synonyms Stereodon cupressiformis Stereodon subimponens
Name authority Hedwig: Sp. Musc. Frond., 291. (1801) Lesquereux: Trans. Amer. Philos. Soc. n. s. 13: 14. (1865)
Web links