Hypnum cupressiforme |
Hypnum bambergeri |
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cypress-leaf claw-moss, cypress-leaf plaitmoss, hypnum moss |
bamberger's hypnum moss, golden claw-moss |
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Habit | Plants small to large, rusty green, golden green, yellow-green, or pale green. | Plants large, reddish to yellowish brown. | ||||||||||||||||
Stem(s) | 1–8+ cm, pale to yellowish green, brown with age, creeping to erect, complanate-foliate or not, irregularly pinnate to nearly unbranched, attached shoots often regularly pinnate, branches 1–3 cm; hyalodermis absent, central strand poorly developed; pseudoparaphyllia filamentous, 1–3-seriate at base. |
leaves falcate to circinate-secund, oblong-ovate to oblong-lanceolate, narrowed to apex, 1.5–2 × 0.4–0.6 mm; base not decurrent, not auriculate; margins plane, sinuate to weakly serrate; acumen slender; costa single, or double and unequal, short to long; alar cells quadrate to short-rectangular, dark brown, region well defined, weakly to markedly excavate, 3–7 cells high along margin, to 3–6 cells wide; basal laminal cells shorter than medial cells, yellow to orange, walls more strongly pitted; medial cells 30–60 × 4–6 µm, walls somewhat to markedly pitted. |
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Leaves | strongly imbricate, not to decidedly falcate-secund, ovate to oblong-lanceolate, gradually or abruptly narrowed to apex, 1.5–2 × 0.5–0.8 mm; base not decurrent, not auriculate; margins recurved to plane proximally, serrulate (sometimes weakly) distally, occasionally nearly entire; acumen slender; costa double and short or obscure; alar region well defined, basalmost cells larger, sometimes hyaline, yellowish or brownish; basal laminal cells shorter, wider than medial cells, not pigmented, walls not pitted; medial cells (50–)60–80 × 3–4(–5) µm. |
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Seta | reddish, 1–2.5(–3) cm. |
reddish, 1.3–2 cm. |
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Sexual condition | dioicous; inner perichaetial leaves oblong-lanceolate, margins serrulate distally, costa obscure. |
dioicous; inner perichaetial leaves ovate to oblong-lanceolate, margins laciniate at shoulders, entire at apex, acumen slender. |
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Capsule | slightly inclined, reddish, cylindric, 1.8–2.5(–2.8) mm; annulus 1–3-seriate; operculum conic to rostrate; endostome cilia 1–2(–3). |
inclined to horizontal, yellowish, oblong-cylindric, 1.3–1.5 mm; annulus 3-seriate; operculum conic; endostome cilia 2 or 3. |
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Hypnum cupressiforme |
Hypnum bambergeri |
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Phenology | Capsules mature Jul–Aug. | |||||||||||||||||
Habitat | Seepage habitats, dry tundra, open conifer forests, mainly calcicolous | |||||||||||||||||
Elevation | low to high elevations (0-2500 m) [low to high elevations (0-8200 ft)] | |||||||||||||||||
Distribution | Nearly worldwide; except Antarctica
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AK; MT; AB; BC; MB; NL; NT; NU; ON; QC; YT; Greenland; Europe; Asia |
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Discussion | Varieties 9 (4 in the flora). Hypnum cupressiforme is an extremely polymorphic species, reflected in the more than 60 varieties that have been described. The species has a wide ecological amplitude as well as a nearly cosmopolitan distribution and is found in all climatic regions except the Antarctic. Taxonomic features reliable in most other species of Hypnum are plastic in H. cupressiforme. Within a single clone, it is possible to sort out several named varieties. Variety lacunosum Bridel was noted by H. Ando (1989) to be weakly differentiated in North America, and he tentatively cited specimens from Nova Scotia, Newfoundland, and South Dakota. Ando noted that var. lacunosum typically is robust, thick-complanate to julaceous, with leaves almost straight to weakly falcate and abruptly narrowed to a short acumen, but the North American material is not robust and is therefore problematic. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Hypnum bambergeri is widely distributed in the Arctic and extends southward, especially in alpine sites. The plants are shiny; the leaf margins curve gradually to the insertion or form a weak auricle just beyond the alar cells. Sporophytes are infrequent. The laminal cells pitted throughout separate this species from all others in Hypnum. The usually calcareous terrestrial seepage habitat is also a useful trait. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 28, p. 537. | FNA vol. 28, p. 536. | ||||||||||||||||
Parent taxa | ||||||||||||||||||
Sibling taxa | ||||||||||||||||||
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Synonyms | Stereodon cupressiformis | Stereodon bambergeri | ||||||||||||||||
Name authority | Hedwig: Sp. Musc. Frond., 291. (1801) | Schimper: Syn. Musc. Eur., 698. (1860) | ||||||||||||||||
Web links |