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Herbs, annual or perennial (stems and leaves glabrous except 43. H. setosum); black glands absent. |
Herbs perennial, semiaquatic or aquatic, erect, branching with long-creeping rhizomes at aerenchymatous base and from mid and distal nodes, 3–10 dm. |
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internodes 4-lined. |
deciduous (base articulated) or persistent (base not articulated). |
ascending to deflexed, sessile; blade narrowly oblong-elliptic (proximal) or lanceolate to linear-lanceolate, 10–30 × 3–8 mm, mostly longer than internodes, not or scarcely smaller distally, leathery, margins plane, apex acute, basal or near-basal veins 1–3(–5), midrib with 0–2 pairs of branches. |
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broadly pyramidal to subcorymbiform, 1(–30)-flowered, branching mostly dichasial. |
3–15 mm diam.; sepals persistent, (4–)5; petals persistent, (4–)5; stamens persistent, (5–)10–80, usually in continuous or interrupted ring, sometimes in 5 barely discernable fascicles, each of 1–2 stamens; ovary (2–)3(–4)-merous; placentation parietal; styles ± spreading, bases distinct; stigmas capitate or clavate. |
4–10 mm diam.; sepals lanceolate, usually unequal, 3–5 × 0.8–1 mm, margins sometimes ciliate, not setulose-ciliate, apex acute to acuminate; petals orange-yellow, obovate, 6–10 mm; stamens 50–80, irregularly grouped; styles 2–4 mm; stigmas capitate. |
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ellipsoid to rostrate-subglobose, 3–4.5 × 2–2.5 mm. |
not carinate. |
0.5–0.6(–0.7) mm; testa obscurely linear-reticulate to irregularly reticulate. |
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= 24. |
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Flowering mid–late summer (Jul–Sep). |
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Open Taxodium swamps, wet pine barrens |
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0–200 m [0–700 ft] |
North America; Mexico; Central America; South America; West Indies; Africa; Asia; Pacific Islands (New Zealand); Australia |
AL; FL; GA; SC |
Species 148 (16 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Herbs annual, wiry; inflorescences: branching mostly monochasial | → 2 |
1. Herbs annual or perennial, not wiry; inflorescences: branching mostly dichasial | → 3 |
2. Leaf blades linear or linear-subulate to linear-lanceolate, 5–22 mm, margins recurved to revolute; sepals 3–7 mm; capsule lengths 1–1.2 times sepals. | H. drummondii |
2. Leaf blades narrowly triangular-subulate to linear-subulate, scalelike, 1–4 mm, margins incurved; sepals 1.5–2.5 mm; capsule lengths 2–2.7 times sepals. | H. gentianoides |
3. Stems and leaves scabrous-tomentose to pilose; sepal margins setulose-ciliate. | H. setosum |
3. Stems and leaves glabrous; sepal margins sometimes ciliate, not setulose-ciliate | → 4 |
4. Leaf blades linear-subulate, 0.2–0.3 mm wide, basal veins 1, midrib unbranched. | H. cumulicola |
4. Leaf blades not linear-subulate, (0.5–)2–18 mm wide, basal veins 1–7, midrib with 0–4 pairs of branches | → 5 |
5. Leaf blades leathery; petals golden yellow or orange-yellow; stamens (35–) 50–80 | → 6 |
5. Leaf blades papery to membranous; petals usually bright, golden, or pale yellow, rarely salmon-orange; stamens 5–25 | → 11 |
6. Leaf blades linear-oblong or linear-lanceolate to linear, 0.5–2 mm wide; petals red tinged; stamens 35–50; styles 2.5–5 mm | H. paucifolium |
6. Leaf blades elliptic, lanceolate, linear-lanceolate, oblong-elliptic, obovate, or ovate, 3–18 mm wide; petals not red tinged; stamens 50–80; styles 2–4 mm | → 7 |
7. Herbs erect or ± spreading, non-aerenchymatous at base; leaves (main stems) longer than internodes | → 8 |
7. Herbs erect to ascending, usually ± aerenchymatous (spongiform-thickened) at base; leaves (main stems) mostly shorter than internodes (usually longer in. 31 H. harperi). | → 9 |
8. Leaf blades (main stem) lanceolate to oblong-elliptic or obovate, 10–30(–50) mm, apex usually acute to acuminate, rarely obtuse; subsidiary inflorescence branches with or without relatively smaller leaves. | H. virgatum |
8. Leaf blades (main stem) narrowly lanceolate, 10–40(–55) mm, apex acute to acuminate; subsidiary inflorescence branches with relatively smaller leaves. | H. radfordiorum |
9. Herbs 2–7 dm; leaf blades (main stem) usually broadly to narrowly ovate, rarely elliptic or lanceolate. | H. denticulatum |
9. Herbs 3–12 dm; leaf blades (main stem) usually lanceolate, sometimes linear-lanceolate or oblong-elliptic, rarely elliptic or ovate | → 10 |
10. Herbs erect to ascending, branching at base and in inflorescence; leaves strongly ascending to appressed, shorter than internodes, smaller distally. | H. erythreae |
10. Herbs erect, branching (from long-creeping rhizomes) at base and from mid and distal nodes; leaves ascending to deflexed, mostly longer than internodes, not or scarcely smaller distally. | H. harperi |
11. Herbs decumbent to ascending, forming loose mats; stigmas scarcely capitate. | H. anagalloides |
11. Herbs usually erect, not forming loose mats; stigmas broadly capitate | → 12 |
12. Leaf blades lanceolate, linear, narrowly elliptic, narrowly oblong-elliptic, oblanceolate, oblanceolate-linear, or obovate; capsules broadest proximal to middle | → 13 |
12. Leaf blades usually elliptic, oblong, broadly oblong-elliptic, ovate, ovate-triangular, round, or suborbiculate, rarely lanceolate; capsules usually broadest at or near middle | → 14 |
13. Leaf blades lanceolate, narrowly oblong-elliptic, or oblanceolate, (2–)6–12 mm wide, basal veins (3–)5–7; inflorescences usually compact. | H. majus |
13. Leaf blades linear to oblanceolate-linear or (proximal) oblanceolate to obovate, 0.5–5.5 mm wide, basal veins 1–3(–5); inflorescences usually diffuse. | H. canadense |
14. Leaf blades (mid and distal) lanceolate-deltate, apex usually subacute; capsules narrowly conic-ellipsoid. | H. gymnanthum |
14. Leaf blades (mid and distal) elliptic, lanceolate, oblanceolate, oblong, round, suborbiculate, or ovate, apex obtuse to rounded; capsules narrowly ovoid to cylindric-ellipsoid | → 15 |
15. Leaf blades bicolor, paler abaxially; inflorescences: bracts linear-subulate. | H. mutilum |
15. Leaf blades concolor; inflorescences: bracts not linear-subulate. | H. boreale |
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FNA vol. 6, p. 88. |
FNA vol. 6, p. 90. Treatment author: Norman K. B. Robson. |
Hypericaceae > Hypericum |
Hypericaceae > Hypericum > sect. Brathys |
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H. adpressum, H. anagalloides, H. apocynifolium, H. ascyron, H. boreale, H. brachyphyllum, H. buckleyi, H. canadense, H. canariense, H. chapmanii, H. cistifolium, H. concinnum, H. crux-andreae, H. cumulicola, H. densiflorum, H. denticulatum, H. dolabriforme, H. drummondii, H. edisonianum, H. ellipticum, H. erythreae, H. fasciculatum, H. frondosum, H. galioides, H. gentianoides, H. graveolens, H. gymnanthum, H. hypericoides, H. kalmianum, H. lissophloeus, H. lloydii, H. lobocarpum, H. maculatum, H. majus, H. microsepalum, H. mutilum, H. myrtifolium, H. nitidum, H. nudiflorum, H. paucifolium, H. perforatum, H. prolificum, H. pseudomaculatum, H. punctatum, H. radfordiorum, H. scouleri, H. setosum, H. sphaerocarpum, H. suffruticosum, H. tenuifolium, H. tetrapetalum, H. virgatum, H. ×mitchellianum |
H. anagalloides, H. boreale, H. canadense, H. cumulicola, H. denticulatum, H. drummondii, H. erythreae, H. gentianoides, H. gymnanthum, H. harperi, H. majus, H. mutilum, H. paucifolium, H. radfordiorum, H. setosum, H. virgatum |
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section Brathys, H. section Brathys, H. section Spachium, H. subsection Spachium, H. section Trigynobrathys, Sarothra section Trigynobrathys |
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unknown |
R. Keller: Bot. Jahrb. Syst. 58: 198. (1923) |
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