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myrtleleaf St. Johnswort

Norton's St. John's-wort, Scouler's St. John's-wort, western john's-wort, western St. John's-wort

Habit Shrubs, erect, with woody caudex, unbranched or branched distally, 3–10 dm, bark on older stems corky. Herbs erect or ascending, with rooting, creeping, branching base, 0.5–6.6(–8) dm.
Stems

internodes 4-lined.

internodes usually weakly 2-lined, sometimes not lined, without black glands, rarely with reddish glands.

Leaves

blades oblong-ovate to triangular-lanceolate, 8–40 × (5–)7–20 mm, base articulated, subcordate-amplexicaul, margins recurved, apex rounded, midrib with 3 or 4 pairs of branches.

usually spreading, rarely erect, sessile or (proximal) subpetiolate;

blade oblong-elliptic or elliptic to triangular-ovate or (proximal) obovate, 12–28(–32) × 6–15(–18) mm, base subcordate to rounded or (proximal) cuneate, margins plane, apex obtuse to rounded, midrib with 4–5 pairs of branches, tertiary veins not densely reticulate, black glands intramarginal (± dense) and, rarely, (1–2) laminar (distal).

Inflorescences

hemispheric to ± flat-topped, 7–30-flowered, widely branched, with flowers or flowering branches from to 3 proximal nodes.

cylindric to narrowly pyramidal, (1–)8–20-flowered.

Flowers

15–25 mm diam.;

sepals persistent, not enclosing capsule, 5, ovate to lanceolate, unequal to subequal, 5–8 × 2–4.5 mm;

petals 5, bright yellow, obovate to oblong-lanceolate, 8–15 mm;

stamens deciduous, 200;

ovary 3(–4)-merous.

6–15(–25) mm diam.;

sepals not or scarcely imbricate, erect in fruit, ovate to lanceolate or narrowly oblong, unequal to subequal, 2.5–5.5 × 1–2 mm, apex acute to rounded;

petals golden yellow, sometimes red-tinged, oblanceolate, 7–12 mm;

stamens 50–90(–109);

anther gland black;

styles 2–8 mm.

Capsules

pyramidal-ovoid, 5–6 × 3–4 mm.

oblanceoloid, 6–10 × 3.5–6 mm, with longitudinal vittae.

Seeds

narrowly carinate, 1 mm;

testa shallowly linear-reticulate.

not carinate, (0.5–)0.7–0.8 mm;

testa linear-reticulate.

2n

= 18.

= 16.

Hypericum myrtifolium

Hypericum scouleri

Phenology Flowering late spring–summer (May–Jul), sometimes fall. Flowering summer (Jun–Sep).
Habitat Moist pinewoods, grassy bogs, pond margins, ditches Wet meadows and banks, coniferous forests, screes, lake margins, marshes, tidal shores
Elevation 0–200 m (0–700 ft) 0–2900 m (0–9500 ft)
Distribution
from FNA
AL; FL; GA; MS; SC
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, Sonora, Zacatecas)
[WildflowerSearch map]
[BONAP county map]
Discussion

Hypericum myrtifolium is related to H. frondosum; it differs in its shorter, usually amplexicaul leaves, the widely dichasially branched inflorescences, and persistent sepals.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Some authors have included Hypericum scouleri in central Mexican H. formosum Kunth as a synonym, subspecies, or variety; the similarities in sepal form and glandularity between these species are due to convergence. Hypericum formosum is related to another Mexican species (H. oaxacanum R. Keller); the affinities of H. scouleri are with the H. attenuatum group from eastern Asia. Hypericum scouleri itself does occur in northern and central Mexico as far south as Michoacán, México, and Hidalgo, where it is known by the synonym H. simulans Rose.

J. M. Gillett and N. K. B. Robson (1981) treated the dwarf alpine form of Hypericum scouleri as subsp. nortoniae. Further work has revealed a range of intermediate forms between the two putative subspecies, which prevents their recognition. Likewise, the differentiation of a southern population (H. formosum subsp. formosum in the sense of C. L. Hitchcock) from a northern one [H. formosum subsp. scouleri (Hooker) C. L. Hitchcock] based on the broader, blunter, and less-glandular sepals in the latter, does not appear warranted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 84. FNA vol. 6, p. 100.
Parent taxa Hypericaceae > Hypericum > sect. Myriandra Hypericaceae > Hypericum > sect. Hypericum
Sibling taxa
H. adpressum, H. anagalloides, H. apocynifolium, H. ascyron, H. boreale, H. brachyphyllum, H. buckleyi, H. canadense, H. canariense, H. chapmanii, H. cistifolium, H. concinnum, H. crux-andreae, H. cumulicola, H. densiflorum, H. denticulatum, H. dolabriforme, H. drummondii, H. edisonianum, H. ellipticum, H. erythreae, H. fasciculatum, H. frondosum, H. galioides, H. gentianoides, H. graveolens, H. gymnanthum, H. harperi, H. hypericoides, H. kalmianum, H. lissophloeus, H. lloydii, H. lobocarpum, H. maculatum, H. majus, H. microsepalum, H. mutilum, H. nitidum, H. nudiflorum, H. paucifolium, H. perforatum, H. prolificum, H. pseudomaculatum, H. punctatum, H. radfordiorum, H. scouleri, H. setosum, H. sphaerocarpum, H. suffruticosum, H. tenuifolium, H. tetrapetalum, H. virgatum, H. ×mitchellianum
H. adpressum, H. anagalloides, H. apocynifolium, H. ascyron, H. boreale, H. brachyphyllum, H. buckleyi, H. canadense, H. canariense, H. chapmanii, H. cistifolium, H. concinnum, H. crux-andreae, H. cumulicola, H. densiflorum, H. denticulatum, H. dolabriforme, H. drummondii, H. edisonianum, H. ellipticum, H. erythreae, H. fasciculatum, H. frondosum, H. galioides, H. gentianoides, H. graveolens, H. gymnanthum, H. harperi, H. hypericoides, H. kalmianum, H. lissophloeus, H. lloydii, H. lobocarpum, H. maculatum, H. majus, H. microsepalum, H. mutilum, H. myrtifolium, H. nitidum, H. nudiflorum, H. paucifolium, H. perforatum, H. prolificum, H. pseudomaculatum, H. punctatum, H. radfordiorum, H. setosum, H. sphaerocarpum, H. suffruticosum, H. tenuifolium, H. tetrapetalum, H. virgatum, H. ×mitchellianum
Synonyms Brathydium myrtifolium, H. glaucum, H. sessiliflorum, Myriandra glauca H. formosum subsp. scouleri, H. formosum var. scouleri, H. nortoniae, H. scouleri subsp. nortoniae
Name authority Lamarck: in J. Lamarck et al., Encycl. 4: 180. (1797) Hooker: Fl. Bor.-Amer. 1: 111. (1831)
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