Hypericum lloydii |
Hypericum scouleri |
|
---|---|---|
sandhill St. Johnswort |
Norton's St. John's-wort, Scouler's St. John's-wort, western john's-wort, western St. John's-wort |
|
Habit | Shrubs, decumbent, straggling and rooting, forming low, rounded clumps or mats, 1–5 dm. | Herbs erect or ascending, with rooting, creeping, branching base, 0.5–6.6(–8) dm. |
Stems | internodes (4-) or 6-lined at first, then terete. |
internodes usually weakly 2-lined, sometimes not lined, without black glands, rarely with reddish glands. |
Leaves | blades linear-subulate, 13–25 × 0.5–0.8 mm, base articulated, parallel, margins revolute, apex rounded to retuse, midrib unbranched. |
usually spreading, rarely erect, sessile or (proximal) subpetiolate; blade oblong-elliptic or elliptic to triangular-ovate or (proximal) obovate, 12–28(–32) × 6–15(–18) mm, base subcordate to rounded or (proximal) cuneate, margins plane, apex obtuse to rounded, midrib with 4–5 pairs of branches, tertiary veins not densely reticulate, black glands intramarginal (± dense) and, rarely, (1–2) laminar (distal). |
Inflorescences | narrowly pyramidal, 1–3-flowered, with 1–3(–5)-flowered dichasia from to 5 proximal nodes, without additional flowering branches; pedicels 0.5 mm. |
cylindric to narrowly pyramidal, (1–)8–20-flowered. |
Flowers | 12–14 mm diam.; sepals deciduous, not enclosing capsule, 5, linear-subulate, unequal, (3–)4.5–7 × 0.5–0.8 mm; petals 5, golden yellow, oblanceolate-oblong, 5–7.5 mm; stamens deciduous, 100; ovary 3-merous. |
6–15(–25) mm diam.; sepals not or scarcely imbricate, erect in fruit, ovate to lanceolate or narrowly oblong, unequal to subequal, 2.5–5.5 × 1–2 mm, apex acute to rounded; petals golden yellow, sometimes red-tinged, oblanceolate, 7–12 mm; stamens 50–90(–109); anther gland black; styles 2–8 mm. |
Capsules | ovoid, 3–4 × 2–2.5 mm. |
oblanceoloid, 6–10 × 3.5–6 mm, with longitudinal vittae. |
Seeds | carinate, 0.7 mm; testa not seen. |
not carinate, (0.5–)0.7–0.8 mm; testa linear-reticulate. |
2n | = 16. |
|
Hypericum lloydii |
Hypericum scouleri |
|
Phenology | Flowering summer (Aug). | Flowering summer (Jun–Sep). |
Habitat | Dry habitats (pine woods, granite outcrops, roadside embankments), inner coastal plain and foothills | Wet meadows and banks, coniferous forests, screes, lake margins, marshes, tidal shores |
Elevation | 100–300 m (300–1000 ft) | 0–2900 m (0–9500 ft) |
Distribution |
AL; GA; NC; SC
|
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, Sonora, Zacatecas)
|
Discussion | The habit, leaf shape, and drier habitats distinguish Hypericum lloydii from H. galioides. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Some authors have included Hypericum scouleri in central Mexican H. formosum Kunth as a synonym, subspecies, or variety; the similarities in sepal form and glandularity between these species are due to convergence. Hypericum formosum is related to another Mexican species (H. oaxacanum R. Keller); the affinities of H. scouleri are with the H. attenuatum group from eastern Asia. Hypericum scouleri itself does occur in northern and central Mexico as far south as Michoacán, México, and Hidalgo, where it is known by the synonym H. simulans Rose. J. M. Gillett and N. K. B. Robson (1981) treated the dwarf alpine form of Hypericum scouleri as subsp. nortoniae. Further work has revealed a range of intermediate forms between the two putative subspecies, which prevents their recognition. Likewise, the differentiation of a southern population (H. formosum subsp. formosum in the sense of C. L. Hitchcock) from a northern one [H. formosum subsp. scouleri (Hooker) C. L. Hitchcock] based on the broader, blunter, and less-glandular sepals in the latter, does not appear warranted. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 79. | FNA vol. 6, p. 100. |
Parent taxa | Hypericaceae > Hypericum > sect. Myriandra | Hypericaceae > Hypericum > sect. Hypericum |
Sibling taxa | ||
Synonyms | H. galioides var. lloydii | H. formosum subsp. scouleri, H. formosum var. scouleri, H. nortoniae, H. scouleri subsp. nortoniae |
Name authority | (Svenson) W. P. Adams: Contr. Gray Herb. 189: 32. (1962) | Hooker: Fl. Bor.-Amer. 1: 111. (1831) |
Web links |
|