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mountain St. Johnswort

millepertuis elliptique, pale Saint John's wort, pale St. John's-wort

Habit Herbs erect, with rooting, creeping base, 3–6.5 dm. Herbs, perennial, erect, with creeping, rhizomatous, ± woody base, usually unbranched, sometimes branched proximally, relatively slender, 1.1–3(–5) dm.
Stems

internodes 4-lined at first, soon 2-lined, with black glands on or near lines.

internodes 4-lined.

Leaves

spreading, sessile;

blade ovate to oblong or lanceolate, 33–65 × 15–27 mm, base cordate to truncate or broadly cuneate, margins plane, apex obtuse to rounded, midrib with 4–5 pairs of branches, tertiary veins densely reticulate toward margins, black glands intramarginal (dense) and, sometimes, laminar (scattered).

blades broadly to narrowly elliptic or oblanceolate to oblong-elliptic, 11–35 × 3–13 mm, base not articulated, cuneate to shallowly cordate-amplexicaul, margins plane to subrevolute, apex rounded, midrib with 5–7 pairs of branches.

Inflorescences

subcorymbiform, (2–)5–14(–22)-flowered, subsidiary branches sometimes narrowly ascending or curved-ascending.

corymbiform, (1–)3–15-flowered, narrowly branched, sometimes with branches from 1–2 proximal nodes.

Flowers

20–25(–30) mm diam.;

sepals not imbricate, erect in fruit, lanceolate, subequal, 5–7.5(–11) × 1–3 mm, apex acute;

petals golden yellow, narrowly obovate, 11–18 mm;

stamens 50–90(–103);

anther gland black;

styles 5.6–12 mm.

12–15 mm diam.;

sepals persistent, not enclosing capsule, (4–)5, ± lanceolate to lanceolate-elliptic, ± unequal, 6–7 × 2–3 mm;

petals (4–)5, pale yellow, sometimes tinged red, obovate to oblanceolate, 6–8 mm;

stamens persistent, 70–95;

ovary 3-merous, placentation parietal.

Capsules

broadly ovoid, 5–8 × 3.5–5 mm, with longitudinal vittae.

ellipsoid to globose, 4–7 × 3.5–5 mm.

Seeds

not carinate, 0.8–1.1 mm;

testa linear-reticulate.

carinate, 0.6–0.7 mm;

testa scalariform-reticulate.

2n

= 16.

= 16, 18.

Hypericum graveolens

Hypericum ellipticum

Phenology Flowering summer (Jun–Aug). Flowering summer (Jun–Sep).
Habitat Open or partly shaded, moist habitats, dry, rocky roadside banks Stream, lake, and pond margins, wet meadows, swamps
Elevation 1200–2100 m (3900–6900 ft) 0–600 m (0–2000 ft)
Distribution
from FNA
NC; TN
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from FNA
CT; IL; MA; MD; ME; MI; MN; NH; NJ; NY; OH; PA; RI; TN; VT; WA; WI; WV; NB; NF; NS; ON; QC
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Discussion

The chromosome count of n = 16 by Adams (in N. K. B. Robson and W. P. Adams 1968) is now regarded as an error; see D. E. Culwell (1970).

Hypericum graveolens is a relict species with close relatives in Japan; it hybridizes with H. ×mitchellianum and, probably, also with H. punctatum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Hypericum ellipticum is related to H. sphaerocarpum, differing by the shorter, herbaceous, rhizomatous habit, shorter leaves, and smaller seeds. A submerged aquatic form (forma submersum Fassett) and one with axillary branches developing after fertilization (forma foliosum Marie-Victorin) seem scarcely worth formal recognition. Hypericum ellipticum is introduced in Washington.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 100. FNA vol. 6, p. 84.
Parent taxa Hypericaceae > Hypericum > sect. Hypericum Hypericaceae > Hypericum > sect. Myriandra
Sibling taxa
H. adpressum, H. anagalloides, H. apocynifolium, H. ascyron, H. boreale, H. brachyphyllum, H. buckleyi, H. canadense, H. canariense, H. chapmanii, H. cistifolium, H. concinnum, H. crux-andreae, H. cumulicola, H. densiflorum, H. denticulatum, H. dolabriforme, H. drummondii, H. edisonianum, H. ellipticum, H. erythreae, H. fasciculatum, H. frondosum, H. galioides, H. gentianoides, H. gymnanthum, H. harperi, H. hypericoides, H. kalmianum, H. lissophloeus, H. lloydii, H. lobocarpum, H. maculatum, H. majus, H. microsepalum, H. mutilum, H. myrtifolium, H. nitidum, H. nudiflorum, H. paucifolium, H. perforatum, H. prolificum, H. pseudomaculatum, H. punctatum, H. radfordiorum, H. scouleri, H. setosum, H. sphaerocarpum, H. suffruticosum, H. tenuifolium, H. tetrapetalum, H. virgatum, H. ×mitchellianum
H. adpressum, H. anagalloides, H. apocynifolium, H. ascyron, H. boreale, H. brachyphyllum, H. buckleyi, H. canadense, H. canariense, H. chapmanii, H. cistifolium, H. concinnum, H. crux-andreae, H. cumulicola, H. densiflorum, H. denticulatum, H. dolabriforme, H. drummondii, H. edisonianum, H. erythreae, H. fasciculatum, H. frondosum, H. galioides, H. gentianoides, H. graveolens, H. gymnanthum, H. harperi, H. hypericoides, H. kalmianum, H. lissophloeus, H. lloydii, H. lobocarpum, H. maculatum, H. majus, H. microsepalum, H. mutilum, H. myrtifolium, H. nitidum, H. nudiflorum, H. paucifolium, H. perforatum, H. prolificum, H. pseudomaculatum, H. punctatum, H. radfordiorum, H. scouleri, H. setosum, H. sphaerocarpum, H. suffruticosum, H. tenuifolium, H. tetrapetalum, H. virgatum, H. ×mitchellianum
Synonyms Brathydium canadense, H. brathydium, H. canadense var. oviforme
Name authority Buckley: Amer. J. Sci. Arts 45: 174. (1843) Hooker: Fl. Bor.-Amer. 1: 110. (1831)
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