Hypericum boreale |
Hypericum harperi |
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millepertuis bor é al, northern bog St. John's-wort, northern St. John's-wort |
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Habit | Herbs annual or perennial, erect, with decumbent to prostrate, rooting base, usually 2–6-branched, branches spreading or ascending distal to middle, 0.9–3.3 dm. | Herbs perennial, semiaquatic or aquatic, erect, branching with long-creeping rhizomes at aerenchymatous base and from mid and distal nodes, 3–10 dm. |
Stems | internodes 4-angled, apical internode shorter than adjacent one or almost absent. |
internodes 4-lined. |
Leaves | spreading, sessile; blade (concolor), broadly to narrowly oblong or elliptic to oblanceolate or round, 5–15 × 2–5 mm, papery to membranous, margins plane, apex rounded, basal veins 3–5, midrib branched or not. |
ascending to deflexed, sessile; blade narrowly oblong-elliptic (proximal) or lanceolate to linear-lanceolate, 10–30 × 3–8 mm, mostly longer than internodes, not or scarcely smaller distally, leathery, margins plane, apex acute, basal or near-basal veins 1–3(–5), midrib with 0–2 pairs of branches. |
Inflorescences | cylindric to rounded-pyramidal, 1–13-flowered, branching mostly dichasial; bracts not linear-subulate. |
broadly pyramidal to subcorymbiform, 1(–30)-flowered, branching mostly dichasial. |
Flowers | 3–5 mm diam.; sepals usually lanceolate to narrowly oblong, rarely oblanceolate, equal, 2.5 × 0.8–1 mm, margins sometimes ciliate, not setulose-ciliate, apex rounded; petals pale yellow, oblong, 1.7–3.5 mm; stamens 5–16, scarcely grouped; styles 0.5 mm; stigmas broadly capitate. |
4–10 mm diam.; sepals lanceolate, usually unequal, 3–5 × 0.8–1 mm, margins sometimes ciliate, not setulose-ciliate, apex acute to acuminate; petals orange-yellow, obovate, 6–10 mm; stamens 50–80, irregularly grouped; styles 2–4 mm; stigmas capitate. |
Capsules | narrowly ovoid to cylindric-ellipsoid, 4–5 × 2–2.5 mm, usually broadest at or near middle. |
ellipsoid to rostrate-subglobose, 3–4.5 × 2–2.5 mm. |
Seeds | 0.4–0.7 mm; testa finely linear-scalariform. |
0.5–0.6(–0.7) mm; testa obscurely linear-reticulate to irregularly reticulate. |
2n | = 16 [“18”]. |
= 24. |
Hypericum boreale |
Hypericum harperi |
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Phenology | Flowering summer–early fall (Jul–Sep). | Flowering mid–late summer (Jul–Sep). |
Habitat | Bogs, poor fens, lake margins, marshes | Open Taxodium swamps, wet pine barrens |
Elevation | 0–500 m (0–1600 ft) | 0–200 m (0–700 ft) |
Distribution |
CT; DE; IA; IN; MA; MD; ME; MI; MN; NH; NJ; NY; OR; PA; RI; VA; VT; WA; WI; BC; MB; NB; NF; NS; ON; PE; QC; SPM |
AL; FL; GA; SC |
Discussion | Where Hypericum boreale grows submerged, the plants are almost always sterile with elongated stems and suborbiculate leaves (H. boreale forma callitrichoides Fassett). Such plants intergrade shorewards with typical H. boreale (F. H. Utech and H. H. Iltis 1970). All other chromosome counts for H. mutilum and its near relatives have given n = 8; B. M. Kapoor’s (1972) count of 2n = 18 must be treated with reserve. Hypericum mutile var. boreale (Britton) E. P. Bicknell is not a validly published name. The discovery of Hypericum boreale near the mouth of Fraser River at Vancouver in 1989 extends the distribution of this species across Canada almost to the Pacific coast; this occurrence is almost certainly the result of recent introduction. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 6, p. 94. | FNA vol. 6, p. 90. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | H. canadense var. boreale, H. mutilum subsp. boreale, Sarothra borealis | |
Name authority | (Britton) E. P. Bicknell: Bull. Torrey Bot. Club 22: 213. (1895) | R. Keller: Bot. Jahrb. Syst. 58: 198. (1923) |
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