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millepertuis bor é al, northern bog St. John's-wort, northern St. John's-wort

Canary Island st john's wort, Canary Islands St. John's wort

Habit Herbs annual or perennial, erect, with decumbent to prostrate, rooting base, usually 2–6-branched, branches spreading or ascending distal to middle, 0.9–3.3 dm. Shrubs erect, bushy, 10–50 dm.
Stems

internodes 4-angled, apical internode shorter than adjacent one or almost absent.

internodes 4-lined at first, then terete.

Leaves

spreading, sessile;

blade (concolor), broadly to narrowly oblong or elliptic to oblanceolate or round, 5–15 × 2–5 mm, papery to membranous, margins plane, apex rounded, basal veins 3–5, midrib branched or not.

spreading, sessile;

blade narrowly elliptic to narrowly elliptic-oblong, 20–70 × 5–15 mm, (proximal usually narrower), base narrowly cuneate to subangustate, margins plane, apex acute to apiculate-obtuse, midrib with 8–12 pairs of branches, tertiary veins densely reticulate toward margins.

Inflorescences

cylindric to rounded-pyramidal, 1–13-flowered, branching mostly dichasial;

bracts not linear-subulate.

broadly rounded-pyramidal to broadly cylindric, to 30-flowered.

Flowers

3–5 mm diam.;

sepals usually lanceolate to narrowly oblong, rarely oblanceolate, equal, 2.5 × 0.8–1 mm, margins sometimes ciliate, not setulose-ciliate, apex rounded;

petals pale yellow, oblong, 1.7–3.5 mm;

stamens 5–16, scarcely grouped;

styles 0.5 mm;

stigmas broadly capitate.

sepals lanceolate, unequal, 3–4.5 × 1–2.2 mm;

petals bright yellow, not red-tinged, oblanceolate-unguiculate, 12–17 mm;

anther gland yellow to orange;

styles widely spreading, 8–14 mm.

Capsules

narrowly ovoid to cylindric-ellipsoid, 4–5 × 2–2.5 mm, usually broadest at or near middle.

pyramidal-ovoid to ovoid-ellipsoid, 9–12 × 7–8 mm.

Seeds

0.4–0.7 mm;

testa finely linear-scalariform.

1.5–2 mm, narrowly winged;

testa linear-reticulate to linear-foveolate.

2n

= 16 [“18”].

= 40.

Hypericum boreale

Hypericum canariense

Phenology Flowering summer–early fall (Jul–Sep). Flowering spring–summer.
Habitat Bogs, poor fens, lake margins, marshes Disturbed sites
Elevation 0–500 m (0–1600 ft) 20–500 m (100–1600 ft)
Distribution
from FNA
CT; DE; IA; IN; MA; MD; ME; MI; MN; NH; NJ; NY; OR; PA; RI; VA; VT; WA; WI; BC; MB; NB; NF; NS; ON; PE; QC; SPM
[BONAP county map]
from FNA
CA; Atlantic Islands (Canary Islands, Madeira) [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Where Hypericum boreale grows submerged, the plants are almost always sterile with elongated stems and suborbiculate leaves (H. boreale forma callitrichoides Fassett). Such plants intergrade shorewards with typical H. boreale (F. H. Utech and H. H. Iltis 1970). All other chromosome counts for H. mutilum and its near relatives have given n = 8; B. M. Kapoor’s (1972) count of 2n = 18 must be treated with reserve.

Hypericum mutile var. boreale (Britton) E. P. Bicknell is not a validly published name.

The discovery of Hypericum boreale near the mouth of Fraser River at Vancouver in 1989 extends the distribution of this species across Canada almost to the Pacific coast; this occurrence is almost certainly the result of recent introduction.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Hypericum canariense is established at Montecito and Santa Barbara in the hills (P. A. Munz 1974) and along the coast north of Santa Cruz to San Francisco, at locations in Orange and San Mateo counties, and in the San Diego coast region.

The description above agrees with that for Hypericum floribundum regarding sepals lanceolate and acute; in typical H. canariense they are oblong-spatulate and rounded. The variation is continuous; only one species is recognized here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 94. FNA vol. 6, p. 96.
Parent taxa Hypericaceae > Hypericum > sect. Brathys Hypericaceae > Hypericum > sect. Webbia
Sibling taxa
H. adpressum, H. anagalloides, H. apocynifolium, H. ascyron, H. brachyphyllum, H. buckleyi, H. canadense, H. canariense, H. chapmanii, H. cistifolium, H. concinnum, H. crux-andreae, H. cumulicola, H. densiflorum, H. denticulatum, H. dolabriforme, H. drummondii, H. edisonianum, H. ellipticum, H. erythreae, H. fasciculatum, H. frondosum, H. galioides, H. gentianoides, H. graveolens, H. gymnanthum, H. harperi, H. hypericoides, H. kalmianum, H. lissophloeus, H. lloydii, H. lobocarpum, H. maculatum, H. majus, H. microsepalum, H. mutilum, H. myrtifolium, H. nitidum, H. nudiflorum, H. paucifolium, H. perforatum, H. prolificum, H. pseudomaculatum, H. punctatum, H. radfordiorum, H. scouleri, H. setosum, H. sphaerocarpum, H. suffruticosum, H. tenuifolium, H. tetrapetalum, H. virgatum, H. ×mitchellianum
H. adpressum, H. anagalloides, H. apocynifolium, H. ascyron, H. boreale, H. brachyphyllum, H. buckleyi, H. canadense, H. chapmanii, H. cistifolium, H. concinnum, H. crux-andreae, H. cumulicola, H. densiflorum, H. denticulatum, H. dolabriforme, H. drummondii, H. edisonianum, H. ellipticum, H. erythreae, H. fasciculatum, H. frondosum, H. galioides, H. gentianoides, H. graveolens, H. gymnanthum, H. harperi, H. hypericoides, H. kalmianum, H. lissophloeus, H. lloydii, H. lobocarpum, H. maculatum, H. majus, H. microsepalum, H. mutilum, H. myrtifolium, H. nitidum, H. nudiflorum, H. paucifolium, H. perforatum, H. prolificum, H. pseudomaculatum, H. punctatum, H. radfordiorum, H. scouleri, H. setosum, H. sphaerocarpum, H. suffruticosum, H. tenuifolium, H. tetrapetalum, H. virgatum, H. ×mitchellianum
Synonyms H. canadense var. boreale, H. mutilum subsp. boreale, Sarothra borealis H. floribundum, Webbia canariensis, W. floribunda
Name authority (Britton) E. P. Bicknell: Bull. Torrey Bot. Club 22: 213. (1895) Linnaeus: Sp. Pl. 2: 784. (1753)
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