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Rusby's rubberweed, Rusby's rubberweed or bitterweed

four-nerve daisy, graylocks four-nerve daisy, graylocks hymenoxys, graylocks rubberweed, old man of the mountain

Habit Perennials, 30–150 cm (polycarpic, often with moderately branched, woody caudices). Perennials, 8–30 cm (polycarpic, often with sparingly branched, woody caudices).
Stems

1–5(–15), usually purple-red-tinted proximally, branched distally, glabrous or ± hairy.

1–10, green throughout to purple-red-tinted proximally or distally to purple-red-tinted throughout, usually unbranched distally, ± hairy.

Leaves

blades simple or lobed (lobes 3), glabrous, gland-dotted;

mid leaves simple or lobed (lobes 3, terminal lobes 2–4 mm wide).

blades simple or lobed (lobes 3–15), ± hairy, gland-dotted (basal leaf bases, sparsely, if at all, long-villous-woolly);

mid leaves simple or lobed (lobes 3–7, terminal lobes 1–2.5 mm wide).

Peduncles

1–3.2 cm, glabrous or ± hairy.

1–10 cm, ± hairy, tomentose distally near involucres.

Involucres

campanulate to urceolate, 6–8 × 6–7 mm.

hemispheric to subglobose, 15–25 × 18–30 mm.

Ray florets

6–8;

corollas yellow, 4.5–8 × 2.5–5 mm.

15–34(–44);

corollas yellow, 16–30 × 4–8 mm.

Disc florets

25–50+;

corollas 2.7–3.2 mm.

150–400+;

corollas 5–6 mm.

Phyllaries

in 2 series, unequal;

outer 7–9, basally connate 1/4–1/3 their lengths, lanceolate, 4.5–6 mm, apices usually rounded;

inner 7–11, obovate, 3.5–4.5 mm, apices mucronate.

in 2–3 series, subequal;

outer 16–24, basally connate 1/5–1/4 their lengths, lanceolate, 9–15 mm, apices acute;

inner 16–24+, lanceolate to oblanceolate, 8–12 mm, apices acuminate to acute.

Heads

50–250+ per plant, in corymbiform arrays.

1–10 per plant, usually borne singly.

Cypselae

narrowly obpyramidal, 2–3 mm;

pappi usually of 5 obovate, sometimes aristate, scales 0.9–1.8 mm.

obpyramidal to narrowly obpyramidal, 3.3–3.7 mm;

pappi of 5–7 lanceolate, aristate scales 4.5–5.3 mm.

2n

= 30.

= 30.

Hymenoxys rusbyi

Hymenoxys grandiflora

Phenology Flowering Jul–Aug(–Oct). Flowering Jun–Aug.
Habitat Roadsides, open areas, edges of forests Meadows, often above timberline
Elevation 1600–2100 m (5200–6900 ft) 2600–4300 m (8500–14100 ft)
Distribution
from FNA
AZ; NM
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; ID; MT; UT; WY
[WildflowerSearch map]
[BONAP county map]
Source FNA vol. 21, p. 443. FNA vol. 21, p. 440.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Gaillardiinae > Hymenoxys Asteraceae > tribe Heliantheae > subtribe Gaillardiinae > Hymenoxys
Sibling taxa
H. ambigens, H. bigelovii, H. brachyactis, H. brandegeei, H. cooperi, H. grandiflora, H. helenioides, H. hoopesii, H. jamesii, H. lemmonii, H. odorata, H. quinquesquamata, H. richardsonii, H. subintegra, H. texana, H. vaseyi
H. ambigens, H. bigelovii, H. brachyactis, H. brandegeei, H. cooperi, H. helenioides, H. hoopesii, H. jamesii, H. lemmonii, H. odorata, H. quinquesquamata, H. richardsonii, H. rusbyi, H. subintegra, H. texana, H. vaseyi
Synonyms Actinella rusbyi Actinella grandiflora, Tetraneuris grandiflora
Name authority (A. Gray) Cockerell: Bull. Torrey Bot. Club 31: 496. (1904) (Torrey & A. Gray) K. F. Parker: Madroño 10: 159. (1950)
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