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hymenopappus, woolly-whites

Habit Biennials or perennials, to 120 cm. Annuals, biennials, or perennials, (5–)20–150 cm.
Stems

erect, often scapiform (1 stem per crown in biennials, usually 3+ in perennials).

Leaves

mostly basal or basal and cauline; alternate;

petiolate;

blades usually 1–2+ times pinnately lobed (lobes usually ± filiform), ultimate margins usually entire, rarely toothed, faces glabrous or hairy, often tomentose, usually gland-dotted as well.

mostly basal or basal and cauline; alternate;

petiolate or sessile;

blades deltate to lanceolate overall, usually 1–2-pinnately or palmati-pinnately lobed, lobes usually filiform, ultimate margins entire or toothed, faces glabrous or hairy, often tomentose, usually gland-dotted.

Involucres

obconic to hemispheric, 4–15+ mm diam.

obconic to hemispheric.

Receptacles

flat or convex, epaleate (except H. newberryi).

flat or convex, usually epaleate (paleae scarious, conduplicate, each ± investing subtended floret in H. newberryi).

Ray florets

0 or 8, pistillate, fertile;

corollas white to ochroleucous.

usually 0, sometimes 8, pistillate, fertile;

corollas white or whitish.

Disc florets

12–70+, bisexual, fertile;

corollas usually yellow to ochroleucous or whitish to purplish, tubes longer than or about equaling funnelform or abruptly dilated, campanulate throats, lobes 5 (reflexed at anthesis).

12–70+, bisexual, fertile;

corollas yellow or whitish to purplish, tubes longer than or about equaling funnelform or abruptly dilated, campanulate throats, lobes 5, ± deltate (reflexed at anthesis);

anther thecae pale;

stigmatic papillae in 2 lines.

Phyllaries

5–13+ in 2–3+ series, subequal, usually membranous- or scarious-margined (often yellowish, whitish, or purplish).

persistent or tardily falling, 5–13+ in 2+ series (distinct, often yellowish, whitish, or purplish, orbiculate to lance-linear, subequal, often ± petaloid, margins usually notably membranous or scarious).

Calyculi

0.

Heads

radiate or discoid, in ± corymbiform arrays.

usually discoid, rarely radiate, usually in ± corymbiform arrays.

Cypselae

obpyramidal (4-, sometimes 5-angled, each face usually 1–4-ribbed, glabrous or hirtellous to villous);

pappi 0 or of 12–22 orbiculate to spatulate scales.

obconic to obpyramidal, usually 4-angled and 12–16-ribbed, glabrous or hairy;

pappi 0 or of 12–22 orbiculate to spatulate scales.

x

= 17.

Hymenopappus

Asteraceae tribe Heliantheae subtribe Hymenopappinae

Distribution
from USDA
North America; Mexico
[BONAP county map]
North America; Mexico H Robinson (1981) suggested that Hymenopappinae is related to Gaillardiinae and other "core" groups of traditional Helenieae
Discussion

Species 11 (10 in the flora).

My treatment of Hymenopappus is essentially an adaptation of that by B. L. Turner (1956).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 5, species 28 (1 genus, 10 species in the flora).

H. Robinson (1981) suggested that Hymenopappinae is related to Gaillardiinae and other “core” groups of traditional Helenieae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Perennials (usually 3+ aerial stems from crown); cauline leaves mostly 0–12; heads 1–20(–50) per stem; disc corolla throat lengths 2–8 times lobes
→ 2
1. Biennials (usually 1 stem from crown); cauline leaves (2–)10–100; heads (15–)20–200 per stem; disc corolla throat lengths 1–2(–3) times lobes
→ 5
2. Ray florets 8
→ 3
2. Ray florets 0
→ 4
3. Basal leaves 12–25 cm; peduncles 6–15 cm; pappi 0, or 0.01–0.1 mm (receptacles paleate)
H. newberryi
3. Basal leaves 8–14 cm; peduncles 3–8 cm; pappi 0.4–0.6 mm
H. radiatus
4. Leaf blades simple or 1-pinnate; cypselae glabrous or sparsely hirtellous
H. mexicanus
4. Leaf blades 2-pinnate; cypselae hirtellous to villous
H. filifolius
5. Ray florets 8
H. biennis
5. Ray florets 0
→ 6
6. Basal leaves simple or 1–2-pinnate, lobes (1–)5–30 mm wide; disc corollas whitish or purplish (not yellowish), tubes 2–3 mm
→ 7
6. Basal leaves 2-pinnate, lobes 0.5–6 mm wide; disc corollas whitish or yellowish, tubes 1.5–2.2 mm
→ 8
7. Basal leaves simple or 1-pinnate; disc corollas usually purplish, rarely whitish; pappi 0.3–2 mm
H. artemisiifolius
7. Basal leaves 1–2-pinnate; disc corollas whitish; pappi 0.1–1 mm
H. scabiosaeus
8. Lobes of basal leaves 1–6 mm wide; disc corollas yellowish; pappi 0.5–1.5 mm
H. flavescens
8. Lobes of basal leaves 0.5–1.5 mm wide; disc corollas whitish; pappi 1–1.5(–2) mm
→ 9
9. Disc corollas 3–4 mm, throats ± funnelform
H. carrizoanus
9. Disc corollas 2.5–3 mm, throats ± campanulate
H. tenuifolius
Source FNA vol. 21, p. 309. Author: John L. Strother. FNA vol. 21, p. 309.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Hymenopappinae Asteraceae > tribe Heliantheae
Subordinate taxa
H. artemisiifolius, H. biennis, H. carrizoanus, H. filifolius, H. flavescens, H. mexicanus, H. newberryi, H. radiatus, H. scabiosaeus, H. tenuifolius
Name authority L’Héritier: Hymenopappus, plate. 1788 Rydberg: in N. L. Britton et al., N. Amer. Fl. 34: 43. (1914)
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