Flora of North America species comparison

appressed-pubescent, trichomes white.

opposite or 3-whorled;

petiole 0.6–2.4 cm, glabrous or sparsely hirsute;

blade elliptic to ovate, (5–)6.4–15 × 3–6(–10) cm, unlobed, base rounded to truncate or cuneate, margins serrate to serrulate, apex usually acute, sometimes acuminate, abaxial surface green, sparsely hirsute, often densely hirsute along veins, trichomes at 40x smooth, 0.5–1 mm, adaxial surface green, glabrate or sparsely hirsute, sometimes moderately hirsute along veins.

usually opposite, sometimes whorled [alternate], simple;

stipules absent;

petiole present or absent;

blade sometimes palmately lobed, margins entire, serrate, serrulate, dentate, denticulate, or crenate;

venation pinnate or acrodromous (Fendlera, Fendlerella, Philadelphus, Whipplea).

open or compact, 250–1000-flowered, usually conic, sometimes ovoid, 7–25 × 6–11.5 cm;

peduncle 2–5.5 cm, sparsely to densely hirsute.

terminal or axillary, cymes, panicles, racemes, or corymbs, or flowers solitary.

1–4 mm, glabrous or sparsely hirsute proximally.

bisexual [unisexual], or sometimes marginal ones sterile, radially symmetric (bisexual ones) or bilaterally symmetric with enlarged petaloid sepals (sterile ones);

perianth and androecium nearly hypogynous, perigynous, or epigynous;

hypanthium completely adnate to ovary or adnate to ovary proximally, free distally;

sepals 4–12, distinct or connate basally;

petals 4–12, connate basally [entirely, then calyptrate];

nectary usually present, rarely absent;

stamens 8–200, usually distinct, sometimes connate proximally, free;

anthers dehiscing by longitudinal slits;

pistil 1, 2–12-carpellate, ovary less than 1/2 inferior, 1/2 inferior, or completely inferior, 1–12-locular, placentation usually axile proximally, parietal distally, rarely strictly axile or parietal;

ovules 1–50 per locule, anatropous;

styles 1–12, distinct or connate proximally to most of length;

stigmas (1–)2–12.

hypanthium adnate to ovary to near its middle, 1–1.8 × 1.2–2.2 mm, weakly 7–10-ribbed in fruit, usually glabrous, rarely sparsely hairy;

sepals shallowly triangular, 0.3–0.7 × 0.5–1.2 mm, margins entire, apex acute to acuminate, abaxial surface glabrous;

petals tardily deciduous, spreading or reflexed, white or yellowish white, elliptic, ovate, or ovate-lanceolate, 3–4 × 1.2–1.5 mm;

filaments 1.8–5 × 0.1–0.3 mm;

anthers 0.3–0.6 mm;

pistil 2–4-carpellate, ovary 1/2 inferior;

styles 2–4, connate to middle or distally, 2–3.5 mm.

capsules [berries], dehiscence septicidal, loculicidal, interstylar, or intercostal.

oblong-ovoid, 3.5–5 × 2–3 mm.

caudate at both ends, 1.7–3 mm.

1–50 per locule, funicular appendage present (Fendlerella, Whipplea) or absent.

flowers present, white or greenish white, sometimes tinged pink or blue, tube 9–16 mm, lobes 4(–5), elliptic to round, (6–)10–21 × (4–)8–15 mm.

= 72, 108 (Japan).