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huperzie brillant, shining clubmoss, shining fir-moss

Chinese clubmoss, fir clubmoss, lycopode de miyoshi, miyoshi's clubmoss, Pacific fir-moss

Gemmiferous branchlets

produced in 1 pseudowhorl at end of each annual growth cycle;

gemmae 4–6 × 3–6 mm, lateral leaves 1.5–2.5 mm wide, broadly obtuse with distinct mucro.

produced in 2–3 pseudowhorls at end of annual growth;

gemmae 3.5–5 × 3–4 mm, lateral leaves 1.25–1.75 mm wide, acute with acuminate tip.

Shoots

erect, indeterminate, 14–20(–100) cm, becoming long-decumbent, with long, trailing, senescent portion turning brown;

juvenile and mature portions similar with strong annual constrictions due to formation of winter bud;

juvenile growth erect.

erect, determinate, 12–18(–25) cm, clustered to long-decumbent, to 8 cm;

leaves in mature portion smaller than leaves in juvenile portion;

annual constrictions absent;

juvenile growth sharply down-curled forming ± 1/2 circle, leaves at tip appressed to form pointed apex.

Leaves

spreading to reflexed, dark green, lustrous;

largest leaves narrowly obovate, leaves broadest at or above middle, 7–11 mm, margins papillate, teeth 1–8, irregular, large;

smallest leaves (at annual constrictions) narrowly lanceolate, 3–6 mm;

stomates on abaxial surface only.

in juvenile (proximal) portion spreading-reflexed (shade) to appressed-ascending (sun), in mature (distal) portion spreading-ascending (shade) to appressed-ascending (sun), light green to yellow, lustrous;

leaves in juvenile portion narrowly lanceolate, parallel-sided, 4.5–7 mm;

leaves in mature portion triangular, widest at base, 3.5–5.5 mm;

margins entire;

stomates present on both surfaces, numerous, 35–80 per 1/2 leaf on adaxial surface.

Spores

23–29 µm. 2n = 134.

25–34 µm.

Huperzia lucidula

Huperzia miyoshiana

Habitat Terrestrial in shaded conifer forests and mixed hardwoods, rarely on rock on shady mossy acidic sandstone On rock or terrestrial on moss-covered boulders in talus slopes, cliffs, near waterfalls, marshes in conifer forest
Elevation 0–1800 m (0–5900 ft) 0–1600(–1800) m (0–5200(–5900) ft)
Distribution
from FNA
AL; AR; CT; DE; GA; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NH; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WI; WV; MB; NB; NF; NS; ON; PE; QC; SPM
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from FNA
AK; WA; BC; Nfld; Asia in Japan; Korea; Siberia
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Discussion

Huperzia × bartleyi (Cusick) Kartesz & Gandhi, a sterile hybrid between H. lucidula and H. porophila, occurs throughout the range of H. porophila and is discussed under that species. Huperzia × buttersii (Abbe) Kartesz & Gandhi is a hybrid between H. lucidula and H. selago.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Lycopodiaceae > Huperzia Lycopodiaceae > Huperzia
Sibling taxa
H. appalachiana, H. haleakalae, H. miyoshiana, H. occidentalis, H. porophila, H. selago
H. appalachiana, H. haleakalae, H. lucidula, H. occidentalis, H. porophila, H. selago
Synonyms Lycopodium lucidulum, Urostachys lucidulus Lycopodium miyoshianum, Lycopodium selago subsp. miyoshianum, Lycopodium selago var. miyoshianum, Lycopodium tenuifolium, Urostachys miyoshiana
Name authority (Michaux) Trevisan: Atti Soc. Ital. Sci. Nat. 17: 248. (1875) (Makino) Ching: Acta Bot. Yunnan. 3(3): 303, 304. (1981)
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