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harlequin lotus, seaside bird's-foot lotus, seaside bird's-foot-trefoil, seaside lotus, witches teeth

balsam bird's-foot trefoil, pine lotus, stipulate lotus, thicket lotus

Habit Herbs usually low, 10–40 cm, glabrous; stoloniferous or rhizomatous. Herbs ± robust, 15–50(–100) cm, usually villous and legume hairy, sometimes glabrous, sometimes glandular-viscid; rhizomatous.
Stems

1–20+, often decumbent, sometimes ascending, unbranched, fleshy, base often spongy.

1–10+, erect to ascending, unbranched or branched, wiry or fleshy.

Leaves

stipules lanceolate-ovate to ovate or deltate, 2–8 mm, ± scarious, fragile; petiolate to long-petiolate;

rachis ± straight, 1–4 cm;

leaflets (3–)5–7, petiolulate, blades elliptic to obovate, 6–20 mm, apex obtuse to emarginate, surfaces glabrous.

stipules widely ovate to lanceolate, (4–)6–9 mm, usually scarious, sometimes leafy, base auriculate or subauriculate; petiolate;

rachis straight, 2–8 cm;

leaflets 9–15(–19), petiolulate to subsessile, blades ovate to obovate, 5–20 mm, apex obtuse to ± acute, surfaces glabrous or hairy, sometimes finely punctate.

Peduncles

ascending to spreading, 2.3–9 cm, equaling or longer than subtending leaf, elongating in fruit, glabrous;

bract subtending umbel, 1–3(–7)-foliolate.

ascending, then spreading, 1–7 cm, shorter than subtending leaf, little elongating in fruit, ± densely hairy, sometimes glandular;

bract medial, (1–)3(–7)-foliolate.

Umbels

(2 or)3–9-flowered.

4–9(or 10)-flowered.

Flowers

10–16 mm;

calyx reddish, 4.5–6 mm, glabrous, tube cylindric, 2.5–4 mm, lobes subulate to narrowly triangular, 2–2.5 mm, eciliate;

corolla banner yellow, wings and keel pale to dark pink, sometimes whitish, wings fading white, keel pink- to purple-tipped, wings ± equaling or longer than keel, claw longer than calyx tube, banner reflexed to 180°, keel apex acute to subacute.

10–12 mm;

calyx green, purplish, or pink, 5–6.5 mm, ± hairy, sometimes glandular, tube cylindric or obconic-cylindric, 4–5 mm, lobes subulate or short-triangular, 1–1.5(–2) mm, ciliolate or eciliate;

corolla banner and keel purple or pink with pink-veined white tips, wings white, longer than keel, claw ± equaling or slightly longer than calyx tube, banner implicate-ascending, keel apex acute.

Legumes

± ascending to ± inclined, brown, oblong, turgid, 20–30 × 2–3 mm, thinly leathery, incompletely septate, glabrous.

± inclined, olive brown to reddish brown, broadly oblong, turgid, 20–25 × (2–)3–4 mm, thinly leathery, incompletely septate, ± hairy to glabrate.

Seeds

(8–)10–14(–16), olive brown to brown, oblong, 1–1.5 mm.

5–7, reddish brown, oblong, to 3 mm.

2n

= 14.

Hosackia gracilis

Hosackia stipularis

Phenology Flowering spring(–early summer).
Habitat Water or springy areas, shores, coastal meadows, wet meadows or woodlands, sphagnum bogs, pastures, roadside ditches, coastal areas.
Elevation 0–200(–700) m. (0–700(–2300) ft.)
Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from USDA
California
[WildflowerSearch map]
[BONAP county map]
Discussion

Hosackia gracilis reaches its northern limit in southeastern Vancouver Island and adjacent Gulf islands near Victoria, British Columbia (G. W. Douglas and M. Ryan 2006). Southward it follows the coast of Washington (Grays Harbor County), Oregon (with an inland incursion to Lane County), and California (to San Luis Obispo County), with disjunct populations inland in the foothills of the Sierra Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems fleshy or wiry; stipules narrow to wide, base sometimes subauriculate.
var. stipularis
1. Stems wiry; stipules wide, base auriculate.
var. ottleyi
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Hosackia Fabaceae > subfam. Faboideae > Hosackia
Sibling taxa
H. alamosana, H. crassifolia, H. incana, H. oblongifolia, H. pinnata, H. rosea, H. stipularis, H. yollabolliensis
H. alamosana, H. crassifolia, H. gracilis, H. incana, H. oblongifolia, H. pinnata, H. rosea, H. yollabolliensis
Subordinate taxa
H. stipularis var. ottleyi, H. stipularis var. stipularis
Synonyms Anisolotus formosissimus, Lotus formosissimus Lotus stipularis
Name authority Bentham: Trans. Linn. Soc. London 17: 365. (1836) Bentham: Trans. Linn. Soc. London 17: 365. (1836)
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